Genus-group names

Pair of Osmia (Osmia) bicornis (Linnaeus, 1758) in precopulation. Foto A. Krebs.

The genus-group names are arranged alphabetically by genus and subgenus, and chronologically within each valid name. Griswold and Michener (1998) and Michener (2007) were used as standard references for the validity, rank and synonymy of genus-group taxa. We deviate from their generic and subgeneric system by keeping Hoplitis (Micreriades) as a distinct subgenus separated from Hoplitis (Alcidamea) and by treating Nasutosmia as a subgenus of Osmia rather than of Hoplitis. Furthermore, Stenosmia and Hoplosmia are reduced to subgeneric rank in Hoplitis and Osmia, respectively, and Pseudoheriades is removed from the osmiine bees. These deviations are justified by the results of a molecular phylogenetic study of the Osmiini (Praz et al., 2008b).  In addition, i) Ceraheriades, a former subgenus of Chelostoma, is treated here as junior synonym of Prochelostoma based on a recent phylogenetic analysis (Sedivy et al., 2008), ii) Exanthocopa, a former subgenus of Hoplitis, is synonymized with the subgenus Anthocopa (A. Müller, unpublished), iii) Ozbekosmia, a former subgenus of Osmia, is synonymized with the subgenus Tergosmia (A. Müller, unpublished), iv) the subgenera Annosmia, Bytinskia, Coloplitis and Hoplitis are merged into one large subgenus Hoplitis based on a recent phylogenetic analysis (Sedivy et al., 2013a, c), v) the subgenera Alcidamea, Megalosmia, Monumetha and Prionohoplitis are merged into one large subgenus Alcidamea based on a recent molecular phylogenetic analysis (Sedivy et al., 2013a, c), and vi) the subgenera Monosmia and Orientosmia are synonymized with the subgenus Osmia based on a combined molecular and morphological phylogeny (Haider et al., 2013).

All alternative published opinions concerning synonymy, rank and generic placement are disclosed citing one reference only (not necessarily the earliest). The following extralimital genera within the osmiine bees restricted to areas outside the Palaearctic are not treated here: Ashmeadiella Cockerell, Atoposmia Cockerell, Bekilia Benoist, Othinosmia Michener and Xeroheriades Griswold. Further, note that several genus-group names are typified by extralimital taxa which are consequently not listed in the following section on species-group names.

Genus Chelostoma Latreille, 1809

1809 Chelostoma Latreille, Genera Crustaceorum et Insectorum […], vol. 4, p. 161 [as genus]. Type species: Apis maxillosa Linnaeus, 1767 [= Chelostoma florisomne (Linnaeus, 1758)], by monotypy.-Synonymy with Heriades Spinola, 1808 in Friese (1911b). Synonymy with Osmia Panzer, 1806 in Warncke (1977-1992).

subgenus Chelostoma Latreille, 1809

1809 Automatically established nominotypical subgeneric taxon (original description and type species are identical with those of genus Chelostoma Latreille.

subgenus Eochelostoma Griswold, 1998

1998 Eochelostoma Griswold, in: Griswold and Michener, Journal of the Kansas Entomological Society, 70: 216 [as subgenus of Chelostoma Latreille]. Type species: Heriades aureocinctus Bingham, 1897 [= Chelostoma aureocinctum (Bingham, 1897)], by original designation.

subgenus Foveosmia Warncke, 1991

1991 Foveosmia Warncke, Linzer Biologische Beiträge, 23: 267 [as subgenus of Osmia Panzer]. Type species: Heriades foveolatus Morawitz, 1868 [= Chelostoma foveolatum (Morawitz, 1868)], by original designation.

subgenus Gyrodromella Michener, 1997

1872 Gyrodroma Thomson, Skandinaviens Hymenoptera, vol. 2, p. 259 [as genus]. NOMEN PRAEOCCUPATUM [not Gyrodroma Klug, 1807 (Hymenoptera)]. Type species: Heriades nigricornis Nylander, 1848 [= Chelostoma rapunculi (Lepeletier, 1841)], by designation of Cockerell (1925: 205).

1997 Gyrodromella Michener, Scientific papers. Natural History Museum. The University of Kansas, 1: 27 [as subgenus of Chelostoma Latreille]. NOMEN NOVUM with same type species for preoccupied Gyrodroma Thomson, 1872 [not Gyrodroma Klug, 1807 (Hymenoptera)].

subgenus Prochelostoma Robertson, 1903

1903 Prochelostoma Robertson, Transactions of the American Entomological Society, 29: 167 [as genus], Type species: Heriades philadelphi Robertson, 1891 [= Chelostoma philadelphi (Robertson, 1891)], by original designation.

1984 Ceraheriades Tkalcu, Annotationes Zoologicae et Botanicae (Bratislava), 158: 5 [as subgenus of Archeriades Peters, now considered to be a synonym of the Afrotropic genus Afroheriades Peters (Michener, 2000: 435)]. Type species: Archeriades petersi Tkalcu, 1984 [= Chelostoma petersi (Tkalcu, 1984)], by original designation.-New synonymy (A. Müller, unpublished).

Genus Haetosmia Popov, 1952

1952 Haetosmia Popov, Trudy Zoologicheskova Instituta, Akademii Nauk SSSR, 10: 104 [as subgenus of Anthocopa Lepeletier & Serville]. Type species: Osmia latipes Morawitz, 1875 [= Haetosmia brachyura (Morawitz, 1875)], by monotypy.-Accepted as subgenus of Osmia Panzer in Peters (1974: 293).

Genus Heriades Spinola, 1808

1808 Heriades Spinola, Insectorium Liguriae […], vol. 2, p. 7 [as genus]. Type species: Apis truncorum Linnaeus, 1758 [= Heriades truncorum (Linnaeus, 1758)], by designation of Latreille (1810: 439).-Synonymy with Osmia Panzer, 1806 in Warncke (1977-1992).

subgenus Heriades Spinola, 1808

1808 Automatically established nominotypical subgeneric taxon (original description and type species are identical with those of genus Heriades Spinola).

1861 Trypetes Schenck, Jahrbücher des Vereins für Naturkunde im Herzogthum Nassau, 14: 89 [as genus]. NOMEN PRAEOCCUPATUM [not Trypetes Schönherr, 1836 (Coleoptera)]. Type species: Apis truncorum Linnaeus, 1758 [= Heriades truncorum (Linnaeus, 1758)], by monotypy.

1895 Eriades Spinola, in: Dalla Torre and Friese, Entomologische Nachrichten (Berlin), 21: 69 [as genus]. Unjustified emendation of Heriades Spinola, 1808.

1987 Orientoheriades Gupta, Reichenbachia (Dresden), 25: 68 [as subgenus of Heriades Spinola]. Type species: Heriades orientalis Gupta, 1987, by original designation, Oriental.-Accepted at genus rank in Gupta (1993: 92).-Synonymy in Michener (2000: 442).

subgenus Michenerella Krombein, 1950

1950 Michenerella Krombein, Proceedings of the Hawaiian Entomological Society, 14: 122 [as subgenus of Heriades Spinola]. Type species: Heriades paganensis Yasumatsu, 1942, by original designation, Oriental.

1955 Eutrypetes Popov, Entomologicheskoe Obozrenie, 34: 280 [as subgenus of Heriades Spinola]. Type species: Heriades turcomanicus Popov, 1955, by original designation.-Synonymy in Michener (2000: 443).

subgenus Rhopaloheriades Griswold & Michener, 1998

1998 Rhopaloheriades Griswold & Michener, Journal of the Kansas Entomological Society, 70: 237 [as subgenus of Heriades Spinola]. Type species: Heriades clavicornis Morawitz, 1875, by original designation.

Genus Hofferia Tkalcu, 1984

1984 Hofferia Tkalcu, Annotationes Zoologicae et Botanicae (Bratislava), 158: 10 [as genus]. Type species: Chelostoma schmiedeknechti Schletterer, 1889 [= Hofferia schmiedeknechti (Schletterer, 1889)], by original designation.

Genus Hoplitis Klug, 1807

1807 Hoplitis Klug, Magazin für Insektenkunde, 6: 225 [as genus]. Type species: Apis adunca Panzer, 1798 [= Hoplitis adunca (Panzer, 1798)], by monotypy.-Synonymy with Osmia Panzer, 1806 in Warncke (1977-1992).

subgenus Alcidamea Cresson, 1864

1864 Alcidamea Cresson, Proceedings of the Entomological Society of Philadelphia, 2: 385 [as genus]. Type species: Alcidamea producta Cresson, 1864 [= Hoplitis producta (Cresson, 1864)], by designation of Michener (1941: 158), Nearctic.

1864 Monumetha Cresson, Proceedings of the Entomological Society of Philadelphia, 2: 387 [as genus]. Type species: Chelostoma albifrons Kirby, 1837 [= Hoplitis argentifrons (Cresson, 1864)], by designation of Titus (1904: 26), Nearctic.-Synonymy in Sedivy et al. (2013c).

1864 Andronicus Cresson, Proceedings of the Entomological Society of Philadelphia, 2: 384 [as genus]. Type species: Andronicus cylindricus Cresson, 1864 (not Giraud, 1857) [= Hoplitis spoliata (Provancher, 1888)], by monotypy, Nearctic.-Synonymy with Monumetha Cresson in Michener (2000: 455).

1872 Acanthosmia Thomson, Skandinaviens Hymenoptera, vol. 2, p. 233 [as subgenus of Osmia Panzer]. No species included, 15 species included in Schmiedeknecht (1885: 20). Type species: Osmia montivaga Morawitz, 1872 [= Hoplitis mitis (Nylander, 1852)], by designation of Sandhouse (1943: 522).-Synonymy in Michener (2000: 449).

1872 Liosmia Thomson, Skandinaviens Hymenoptera, vol. 2, p. 233, p. 254 [as subgenus of Osmia Panzer]. No species included, seven species included in Zanden (1988b: 122). Type species: Osmia claviventris Thomson, 1872 [= Hoplitis claviventris (Thomson, 1872)], by designation of Michener (1941: 159).-Synonymy in Michener (2000: 449).

1885 Tridentosmia Schmiedeknecht, Apidae Europaeae, vol. 2, p. 21 [887] [as subgenus of Osmia Panzer]. Type species: Osmia tridentata Dufour & Perris, 1840 [= Hoplitis tridentata (Dufour & Perris, 1840)], by designation of Michener (1941: 159).-Synonymy in Michener (2000: 449).

1885 Megalosmia Schmiedeknecht, Apidae Europaeae, vol. 2, p. 23 [889] [as subgenus of Osmia Panzer]. Type species: Osmia grandis Morawitz, 1872 [= Hoplitis fulva (Eversmann, 1852)], by designation of Sandhouse (1943: 568).-Synonymy in Sedivy et al. (2013c).

1916 Autochelostoma Sladen, The Canadian Entomologist, 48: 270 [as genus]. Type species: Autochelostoma canadensis Sladen, 1916 [= Hoplitis producta (Cresson, 1864)], by original designation and monotypy.-Synonymy in Michener (2000: 449).

1916 Chlorosmia Sladen, The Canadian Entomologist, 48: 270 [as genus]. Type species: Osmia fulgida Cresson, 1864 [= Hoplitis fulgida (Cresson, 1864)], by monotypy, Nearctic.-Synonymy with Monumetha Cresson in Michener (2000: 455).

1993 Prionohoplitis Tkalcu, Bulletin de la Société Entomologique de Mulhouse, July-September: 55 [as subgenus of Hoplitis Klug]. Type species: Osmia curvipes Morawitz, 1871 [= Hoplitis curvipes (Morawitz, 1871)], by original designation.-Synonymy in Sedivy et al. (2013c).

1995 Calohoplitis Tkalcu, Entomologische Abhandlungen (Dresden), 57: 129 [as subgenus of Hoplitis Klug]. Type species: Osmia laboriosa Smith, 1878 [= Hoplitis laboriosa (Smith, 1878)], by original designation.-Synonymy with Megalosmia Schmiedeknecht in Michener (2000: 454)

subgenus Anthocopa Lepeletier & Serville, 1825

1825 Anthocopa Lepeletier & Serville, Encyclopédie Méthodique, Histoire Naturelle. Insectes, vol. 10, p. 314 [as genus]. Type species: Apis papaveris Latreille, 1799 [= Hoplitis papaveris (Latreille, 1811)], by monotypy.

1860 Phyllotoma Duméril, Mémoires de l’Académie des Sciences de l’Institut de France (Paris), 31: 842 [as genus]. NOMEN PRAEOCCUPATUM [not Phyllotoma Leach, 1819 (Coleoptera); not Phyllotoma Fallén, 1829 (Hymenoptera)]. Type species: Apis papaveris Latreille, 1799 [= Hoplitis papaveris (Latreille, 1799)], by designation of Michener (1941: 160).

1872 Pseudosmia Radoszkowski, Horae Societatis Entomologicae Rossicae (St. Petersburg), 8, Bulletin: xviii [as genus]. No species included, nine species included in Radoszkowski (1874c: 152). Type species: Megachile cristata Fonscolombe, 1846 (not Dufour, 1841) [= Hoplitis cristatula (Zanden, 1990)], by designation of Cockerell (1922: 6).-Synonymy in Michener (2000: 450).

1874 Pseudo-Osmia Radoszkowski, Bulletin de la Société Impériale des Naturalistes de Moscou, 46[47]: 137 [as genus]. UNJUSTIFIED EMENDATION of Pseudosmia Radoszkowski, 1872.

1874 Pseudoosmia Radoszkowski, Bulletin de la Société Impériale des Naturalistes de Moscou, 48: 152 [as genus]. UNJUSTIFIED EMENDATION of Pseudosmia Radoszkowski, 1872.

1885 Arctosmia Schmiedeknecht, Apidae Europaeae, vol. 2, p. 21 [887] [as subgenus of Osmia Panzer]. Type species: Megachile villosa Schenck, 1853 (not Fabricius, 1775) [= Hoplitis villosa (Schenck, 1853)], by monotypy.-Synonymy in Michener (2000: 450).

1885 Furcosmia Schmiedeknecht, Apidae Europaeae, vol. 2, p. 22 [888] [as subgenus of Osmia Panzer]. Type species: Apis papaveris Latreille, 1799 [= Hoplitis papaveris (Latreille, 1799)], by designation of Cockerell (1922: 6).-Synonymy in Michener (2000: 450).

1886 Pseudocosmia Radoszkowski, Horae Societatis Entomologicae Rossicae (St. Petersburg), 20: 14 [as genus]. UNJUSTIFIED EMENDATION of Pseudosmia Radoszkowski, 1872.

1935 Lithosmia Alfken, Veröffentlichungen aus dem Deutschen Kolonial- und Übersee-Museum in Bremen, 1: 188 [as subgenus of Osmia Panzer]. Type species: Megachile villosa Schenck, 1853 (not Fabricius, 1775) [= Hoplitis villosa (Schenck, 1853)], by designation and synonymization with subgenus Arctosmia Schmiedeknecht in Sandhouse (1943: 565).-Synonymy with subgenus Arctosmia Schmiedeknecht previously in Alfken (1935c: 314). Synonymy in Michener (2000: 442).

1943 Glossosmia Michener, Annals of the Entomological Society of America, 36: 84 [as subgenus of Osmia Panzer]. Type species: Osmia singularis Morawitz, 1875 [= Hoplitis singularis (Morawitz, 1875)], by original designation.-Synonymy in Michener (2000: 450).

1993 Exanthocopa Tkalcu, Bulletin de la Société Entomologique de Mulhouse, July-September: 55 [as subgenus of Anthocopa Lepeletier & Serville]. Type speciesOsmia oxypyga Benoist, 1927 [= Hoplitis oxypyga (Benoist, 1927)], by original designation.-New synonymy (A. Müller, unpublished).

subgenus Chlidoplitis Griswold, 1998

1998 Chlidoplitis Griswold, in: Griswold and Michener, Journal of the Kansas Entomological Society, 70: 223 [as subgenus of Hoplitis Klug]. Type species: Hoplitis heinrichi Zanden, 1980, by original designation.

subgenus Eurypariella Tkalcu, 1995

1995 Eurypariella Tkalcu, Entomologische Abhandlungen (Dresden), 57: 111 [as genus]. Type species: Osmia denudata Morawitz, 1880 [= Hoplitis denudata (Morawitz, 1880)], by original designation.

subgenus Formicapis Sladen, 1916

1916 Formicapis Sladen, The Canadian Entomologist, 48: 271 [as genus]. Type species: Formicapis clypeata Sladen, 1916 [= Hoplitis robusta (Nylander, 1848)], by monotypy.

subgenus Hoplitis Klug, 1807

1807 Automatically established nominotypical subgeneric taxon (original description and type species are identical with those of genus Hoplitis Klug).

1872 Ctenosmia Thomson, Skandinaviens Hymenoptera, vol. 2, p. 233 [as subgenus of Osmia Panzer]. Type species: Apis adunca Panzer, 1798 [= Hoplitis adunca (Panzer, 1798)], by designation and synonymization in Michener (1941: 158).-Synonymy in Michener (2000: 453).

1954 BytinskiaMavromoustakis, Bulletin of the Research Council of Israel, 4: 269 [as genus]. Type species: Bytinskia erythrogastra Mavromoustakis, 1954 [= Hoplitis erythrogastra (Mavromoustakis, 1954)], by original designation.-Synonymy in Sedivy et al. (2013c).

1991 Annosmia Warncke, Linzer Biologische Beiträge, 23: 307 [as subgenus of Osmia Panzer]. Type species: Osmia annulata Latreille, 1811 [= Hoplitis annulata (Latreille, 1811)], by original designation.-Synonymy in Sedivy et al. (2013c).

1998 Coloplitis Griswold, in: Griswold and Michener, Journal of the Kansas Entomological Society, 70: 223 [as subgenus of Hoplitis Klug]. Type species: Hoplitis premordica Griswold, 1998, by original designation.-Synonymy in Sedivy et al. (2013c).

subgenus Jaxartinula Popov, 1963

1963 Jaxartinula Popov, Zoologicheskiy Zhurnal (Moscow), 42: 865 [as genus]. Type species: Jaxartinula malyshevi Popov, 1963 [= Hoplitis malyshevi (Popov, 1963) ], by original designation.

subgenus Kumobia Popov, 1962

1962 Kumobia Popov, Trudy Zoologiceskzo Instituta Akademija Nauk SSSR, 30: 300 [as genus]. Type species: Osmia tenuicornis Morawitz, 1875 [= Hoplitis tenuicornis (Morawitz, 1875)], by original designation.

subgenus Megahoplitis Tkalcu, 1993

1993 Megahoplitis Tkalcu, Bulletin de la Société Entomologique de Mulhouse, July-September: 55 [as subgenus of Hoplitis Klug]. Type species: Osmia tigrina Morawitz, 1872 [= Hoplitis tigrina (Morawitz, 1872)], by original designation.

subgenus Micreriades Mavromoustakis, 1958

1958 Micreriades Mavromoustakis, The Annals and Magazine of Natural History, ser. 13, 1: 444 [as subgenus of Heriades Spinola]. Type species: Heriades parnesicus Mavromoustakis, 1958 [= Hoplitis parnesica (Mavromoustakis, 1958)], by original designation.-Synonymy with subgenus Alcidamea Cresson, 1864 in Michener (2000: 449).

subgenus Pentadentosmia Warncke, 1991

1991 Pentadentosmia Warncke, Entomofauna (Ansfelden), 12: 14 [as subgenus of Osmia Panzer]. Type species: Osmia quinquespinosa Friese, 1899 [= Hoplitis quinquespinosa (Friese, 1899)], by original designation.

subgenus Platosmia Warncke, 1990

1990 Platosmia Warncke, Entomofauna (Ansfelden), 11: 482 [as subgenus of Osmia Panzer]. Type species: Osmia platalea Warncke, 1990 [= Hoplitis platalea (Warncke, 1990)], by original designation.

subgenus Stenosmia Michener, 1941

1941 Stenosmia Michener, American Midland Naturalist, 26: 165 [as genus]. Type species: Hoplitis crassipunctata Müller, 2014, newly fixed due to the misidentification of Hoplitis flavicornis (Morawitz, 1877) in the original designation by Michener (1941).

subgenus Tkalcua Kocak & Kemal, 2010

1993 Microhoplitis Tkalcu, Veröffentlichungen des Übersee-Museum Bremen (Naturwissenschaften), 12: 811 [as subgenus of Osmia Panzer]. NOMEN PRAEOCCUPATUM [not Microhoplitis Marumo, 1920 (Lepidoptera)]. Type species: Osmia hohmanni Tkalcu, 1993 [= Hoplitis zandeni (Teunissen & van Achterberg, 1992)], by original designation.

2010 Tkalcua Kocak & Kemal, Priamus, 12: 128 [as subgenus of Hoplitis Klug]. NOMEN NOVUM with same type species for preoccupied Microhoplitis Tkalcu, 1993 [not Microhoplitis Marumo, 1920 (Lepidoptera)].

Genus Ochreriades Mavromoustakis, 1956

1956 Ochreriades Mavromoustakis, Eos (Madrid), 32: 226 [as genus]. Type species: Heriades fasciatus Friese, 1899 [= Ochreriades fasciatus (Friese, 1899)], by original designation.

Genus Osmia Panzer, 1806

1806 Osmia Panzer, Apis Calendarum, Hymenoptera 2, vol. 6, p. 230 [as genus]. Type species: Apis bicornis Linnaeus, 1758 [= Osmia bicornis (Linnaeus, 1758)], by designation of Latreille (1810: 439).

subgenus Allosmia Tkalcu, 1974

1974 Allosmia Tkalcu, Beiträge zur Entomologie (Berlin), 24: 331 [as subgenus of Osmia Panzer]. Type species: Osmia rufohirta Latreille, 1811, by original designation.

subgenus Erythrosmia Schmiedeknecht, 1885

1885 Erythrosmia Schmiedeknecht, Apidae Europaeae, vol. 2, p. 20 [886] [as subgenus of Osmia Panzer]. Type species: Osmia andrenoides Spinola, 1808, by designation of Cockerell (1922: 6).

subgenus Helicosmia Thomson, 1872

1872 Helicosmia Thomson, Skandinaviens Hymenoptera, vol. 2, p. 233 [as subgenus of Osmia Panzer]. No species included, seven species included in Schmiedeknecht (1885: 22). Type species: Apis aurulenta Panzer, 1799 [= Osmia aurulenta (Panzer, 1799)], by designation of Michener (1941: 163).

1885 Chalcosmia Schmiedeknecht, Apidae Europaeae, vol. 2, p. 20 [886] [as subgenus of Osmia Panzer]. Type species: Apis fulviventris Panzer, 1798 (not Scopoli, 1763) [= Osmia niveata (Fabricius, 1804)], by designation of Sandhouse (1939: 13).-Synonymy in Michener (2000: 464).

1903 Gnathosmia Robertson, Transactions of the American Entomological Society, 29: 165 [as genus]. Type species: Osmia georgica Cresson, 1878, by original designation and monotypy, Nearctic.-Synonymy with subgenus Chalcosmia Schmiedeknecht, 1885 in Sandhouse (1939: 13).-Synonymy in Michener (2000: 464).

1950 Cryptosmia Yasumatsu & Hirashima, Mushi (Fukuoka), 21: 14 [as subgenus of Osmia Panzer]. Type species: Osmia satoi Yasumatsu & Hirashima, 1950, by original designation.-Synonymy in Michener (2000: 464).

subgenus Hemiosmia Tkalcu, 1975

1975 Hemiosmia Tkalcu, Acta Entomologica Bohemoslovaca (Praha), 72: 34 [as subgenus of Osmia Panzer]. Type species: Osmia argyropyga Pérez, 1879, by original designation.

1979 Exosmia Tkalcu, Acta Entomologica Bohemoslovaca (Praha), 76: 321 [as subgenus of Osmia Panzer]. Type species: Osmia difficilis Morawitz, 1875, by original designation.-Synonymy in Michener (2000: 464).

subgenus Hoplosmia Thomson, 1872

1872 Hoplosmia Thomson, Skandinaviens Hymenoptera, vol. 2, p. 233 [as subgenus of Osmia Panzer]. No species included, six species included in Schmiedeknecht (1885: 23). Type species: Apis spinulosa Kirby, 1802 [= Osmia spinulosa (Kirby, 1802)], by designation of Michener (1941: 161).

1974 Odontanthocopa Tkalcu, Acta Entomologica Bohemoslovaca (Praha), 71: 125 [as subgenus of Anthocopa Lepeletier & Serville]. Type species: Osmia bidentata Morawitz, 1876, by original designation.-New synonymy (A. Müller, unpublished).

1974 Odonterythrosmia Tkalcu, Acta Entomologica Bohemoslovaca (Praha), 71: 131 [as subgenus of Anthocopa Lepeletier & Serville]. Type species: Osmia fallax Pérez, 1895, by original designation.-Synonymy with Odontanthocopa in Michener (2000: 458).

1974 Paranthocopa Tkalcu, Acta Entomologica Bohemoslovaca (Praha), 71: 132 [as subgenus of Anthocopa Lepeletier & Serville]. Type species: Osmia pinguis Pérez, 1895, by original designation.-New synonymy (A. Müller, unpublished).

subgenus Melanosmia Schmiedeknecht, 1885

1885 Melanosmia Schmiedeknecht, Apidae Europaeae, vol. 2, p. 19 [885] [as subgenus of Osmia Panzer]. Type species: Osmia fuciformis Latreille, 1811 [= Osmia xanthomelana (Kirby, 1802)], by designation of Sandhouse (1939: 33).

1899 Nothosmia Ashmead, Transactions of the American Entomological Society, 26: 75 [as genus]. Type species: Osmia distincta Cresson, 1864, by original designation, Nearctic.-Synonymy in Michener (2000: 465).

1903 Centrosomia Robertson, Transactions of the American Entomological Society, 29: 165, 166 [as genus]. Type species: Osmia bucephala Cresson, 1864, by original designation and monotypy, Nearctic.-Synonymy in Michener (2000: 465).

1903 Leucosmia Robertson, Transactions of the American Entomological Society, 29: 166, 171 [as genus]. Type species: Osmia albiventris Cresson, 1864, by original designation and monotypy, Nearctic.-Synonymy with subgenus Nothosmia Ashmead, 1899 in Hurd (1979: 2039). Synonymy in Michener (2000: 465).

1903 Monilosmia Robertson, Transactions of the American Entomological Society, 29, 166, 171 [as genus]. Type species: Osmia canadensis Cresson, 1864 [= Osmia simillima Smith, 1853], by original designation and monotypy, Nearctic.-Synonymy in Michener (2000: 465).

1903 Xanthosmia Robertson, Transactions of the American Entomological Society, 29: 166, 171 [as genus]. Type species: Osmia cordata Robertson, 1902, by original designation and monotypy, Nearctic.-Synonymy with subgenus Nothosmia Ashmead, 1899 in Hurd (1979: 2039). Synonymy in Michener (2000: 465).

1958 Chenosmia Sinha, University of Kansas Science Bulletin, 39: 233 [as genus]. Type species: Osmia penstemonis Cockerell, 1906, by original designation, Nearctic.-Synonymy in Michener (2000: 465).

subgenus Metallinella Tkalcu, 1966

1966 Metallinella Tkalcu, Acta Entomologica Bohemoslovaca (Praha), 63: 200 [as genus]. Type species: Osmia atrocaerulea Schilling, 1849 [= Osmia brevicornis (Fabricius 1798)], by original designation.-Synonymy with Nearctic subgenus Cephalosmia Sladen, 1916 in Warncke (1991d: 283).

subgenus Nasutosmia Griswold & Michener, 1998

1998 Nasutosmia Griswold & Michener, Journal of the Kansas Entomological Society, 70: 225 [as subgenus of Hoplitis Klug]. Type species: Osmia nasuta Friese, 1899, by original designation.

subgenus Neosmia Tkalcu, 1974

1974 Neosmia Tkalcu, Beiträge zur Entomologie (Berlin), 24: 333 [as subgenus of Osmia Panzer]. Type species: Osmia gracilicornis Pérez, 1895, by original designation.

subgenus Osmia Panzer, 1806

1806 Automatically established nominotypical subgeneric taxon (original description and type species are identical with those of genus Osmia Panzer).

1807 Amblys Klug, Magazin für Insektenkunde, 6: 198 [as genus]. Type species: Apis bicornis Linnaeus, 1758 [= Osmia bicornis (Linnaeus, 1758)], by designation of Latreille (1811: 577).-Synonymy in Michener (2000: 467).

1872 Ceratosmia Thomson, Skandinaviens Hymenoptera, vol. 2, p. 232 [as subgenus of Osmia Panzer]. Type species: Apis bicornis Linnaeus, 1758 [= Osmia bicornis (Linnaeus, 1758)], by designation of Sandhouse (1939: 9).-Synonymy in Michener (2000: 467).

1885 Aceratosmia Schmiedeknecht, Apidae Europaeae, vol. 2, p. 19 [885] [as subgenus of Osmia Panzer]. Type species: Osmia emarginata Lepeletier, 1841, by designation of Sandhouse (1939: 9).-Synonymy in Michener (2000: 467).

1900 Pachyosmia Ducke, Bericht des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck, 25: 18 [as subgenus of Osmia Panzer]. Type species: Apis rufa Linnaeus, 1758 [= Osmia bicornis (Linnaeus, 1758)], by designation of Sandhouse (1939: 9).-Synonymy in Michener (2000: 467).

1974 Monosmia Tkalcu, Beiträge zur Entomologie (Berlin), 24: 337 [as subgenus of Osmia Panzer]. Type speciesOsmia apicata Smith, 1853, by original designation.-Synonymy in Haider et al. (2013).

1978 Orientosmia Peters, Senckenbergiana Biologica (Frankfurt), 58: 332 [as subgenus of Osmia Panzer]. Type speciesOsmia maxillaris Morawitz, 1875, by original designation.-Synonymy in Haider et al. (2013).

subgenus Pyrosmia Tkalcu, 1975

1975 Pyrosmia Tkalcu, Acta Rerum Naturalium Musei Nationalis Slovaci Bratislava, 21: 182 [as subgenus of Osmia Panzer]. Type species: Osmia ferruginea Latreille, 1811, by original designation.

1988 Caerulosmia Zanden, Zoologische Mededelingen (Leiden), 62: 123 [as subgenus of Osmia Panzer]. NOMEN NUDUM according to Michener (2000: 468).

1988 Viridosmia Warncke, Entomofauna (Ansfelden), 9: 5 [as subgenus of Osmia Panzer]. Type species: Osmia saxicola Ducke, 1899, by original designation.-Synonymy in Michener (2000: 468).

1989 Caerulosmia Zanden, Entomologische Abhandlungen (Dresden), 53: 83 [as subgenus of Osmia Panzer]. Type species: Osmia gallarum Spinola, 1808, by original designation.-Synonymy in Michener (2000: 468).

subgenus Tergosmia Warncke, 1988

1988 Tergosmia Warncke, Entomofauna (Ansfelden), 9: 390 [as subgenus of Osmia Panzer]. Type species: Osmia tergestensis Ducke, 1897, by original designation.

1993 Heterosmia Tkalcu, Bulletin de la Société Entomologique de Mulhouse, July-September: 56 [as subgenus of Osmia Panzer]. NOMEN NUDUM according to Michener (2000: 468).

1994 Ozbekosmia Zanden, Reichenbachia (Dresden), 30: 168 [as subgenus of Osmia Panzer]. Type species: Osmia avosetta Warncke, 1988, by original designation.-New synonymy (A. Müller, unpublished).

Genus Protosmia Ducke, 1900

1900 Protosmia Ducke, Bericht des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck, 25: 12 [as subgenus of Osmia Panzer]. Type species: Heriades glutinosus Giraud, 1871 [= Protosmia glutinosa (Giraud, 1871)], by designation of Michener (1943: 86) and Sandhouse (1943: 592).

subgenus Chelostomopsis Cockerell, 1925

1925 Chelostomopsis Cockerell, Proceedings of the California Academy of Sciences, 14: 205 [as genus]. Type species: Chelynia rubifloris Cockerell, 1898 [= Protosmia rubifloris (Cockerell, 1898)], by original designation, Nearctic.

1936 Raphidostoma Cockerell, The Pan-Pacific Entomologist, 12: 133 [as genus]. Type species: Raphidostoma ceanothi Cockerell, 1936 [= Protosmia rubifloris (Cockerell, 1898)], by original designation, Nearctic.-Synonymy in Michener (2000: 470).

subgenus Nanosmia Griswold, 1998

1998 Nanosmia Griswold, in: Griswold and Michener, Journal of the Kansas Entomological Society, 70: 241 [as subgenus of Protosmia Ducke]. Type species: Protosmia asensioi Griswold & Parker, 1987, by original designation.

subgenus Protosmia Ducke, 1900

1900 Automatically established nominotypical subgeneric taxon (original description and type species are identical with those of genus Protosmia Ducke).

1943 Rhodosmia Michener, Annals of the Entomological Society of America, 36: 85 [as subgenus of Anthocopa Lepeletier & Serville]. Type species: Osmia paradoxa Friese, 1899 [= Protosmia paradoxa (Friese, 1899)], by original designation.-Synonymy in Michener (2000: 471).

Genus Stenoheriades Tkalcu, 1984

1984 Stenoheriades Tkalcu, Annotationes Zoologicae et Botanicae (Bratislava), 158: 1 [as subgenus of Pseudoheriades Peters]. Type species: Pseudoheriades hofferi Tkalcu, 1984 [= Stenoheriades coelostoma (Benoist, 1935)], by original designation.

Genus Wainia Tkalcu, 1980

1980 Wainia Tkalcu, Annotationes Zoologicae et Botanicae (Bratislava), 135: 1 [as genus]. Type species: Wainia lonavlae Tkalcu, 1980, by original designation, Oriental.

subgenus Caposmia Peters, 1984

1984 Caposmia Peters, Entomofauna (Ansfelden), 5: 378 [as subgenus of Osmia Panzer]. Type species: Osmia braunsi Peters, 1984 [= Wainia braunsi (Peters, 1984)], by original designation, Afrotropic.

1991 Eremoplosmia Zanden, Reichenbachia (Dresden), 28: 164 [as subgenus of Anthocopa Lepeletier & Serville]. Type species: Osmia eremoplana Mavromoustakis, 1949 [= Wainia eremoplana (Mavromoustakis, 1949)], by original designation.-Accepted as subgenus of Osmia Panzer in Griswold and Michener (1998: 229, 231, 234) by mistake [see their erratum].-Synonymy in Michener (2000: 473).

subgenus Wainiella Griswold, 1998

1998 Wainiella Griswold, Journal of the Kansas Entomological Society, 70: 234 [as subgenus of Wainia Tkalcu]. Type species: Heriades sakaniensis Cockerell, 1936 [= Wainia sakaniensis (Cockerell, 1936)], by original designation, Afrotropic.