Palaearctic Osmiine Bees

Female of Osmia (Osmia) bicornis bicornis (Linnaeus, 1758) collecting moist soil for the construction of the brood cells. Foto N. Vereecken.

Listed below are three extinct osmiine taxa described from their fossilized remains in the Palaearctic region. Further compression fossils of putative Osmiini taxa have been described from Florissant, Colorado, USA (Eocene-Oligocene) (see Zeuner and Manning, 1976 and Engel and Perkovsky, 2006 for reviews). The Palaearctic Baltic amber fossils described in detail by Engel (2001) on the other hand are attributed to the now extinct tribes Ctenoplectrellini Engel and Glyptapini Cockerell (Engel and Perkovsky, 2006) even though they had been classified as subtribes (Ctenoplectrellina, Glyptapina) within the tribe Osmiini before (Engel, 2001). As the correct tribal, let alone generic identification is often doubtful due to the fragmentary evidence, these taxa are listed under Incertae sedis.

† Osmia antiqua Heer, 1849

1849 † Osmia antiqua Heer, Die Insektenfauna der Tertiärgebilde von Oeningen und von Radoboj in Croatien, vol. 2, p. 95. Compression fossil. Upper Miocene. Type material: Holotype Oe. 91, Inv. 1272, “Oeningen” [Germany], SMNK (Karlsruhe).

† Osmia carbonum Heyden, 1862

1862 † Osmia carbonum Heyden, Palaeontographica, 10: 75. Compression fossil. Upper Oligocene. Type material: Holotype 58779, “Rott im Siebengebirge” [Germany], BMNH (London); paratype 58768.

† Osmia nigra Zeuner & Manning, 1976

1976 † Osmia nigra Zeuner & Manning, Bulletin of the British Museum (Natural History), Geology, 27: 185. Compression fossil. Upper Miocene. Type material: Holotype, Pl. I. 719, “Oeningen” [Germany], ETHZ (Zürich); paratype, Pl. I. 685.

Incertae sedis

Family-group names

2001 † Ctenoplectrellina Engel, 2001, Bulletin of the American Museum of Natural history, 259: 54. Type genus: Ctenoplectrella Cockerell, 1909.

Notes-Engel (2001: 54-55) proposed this taxon as subtribe within the Osmiini, but indicated that a future raise of its rank to tribal level was likely and in Engel (2005: 32) and Engel and Perkovsky (2006: 6) it was indeed accepted as tribe as presented in our Table 2.

1909 † Glyptapina Cockerell, Schriften der physikalisch-ökonomischen Gesellschaft zu Königsberg, 50: 13. Type genus: Glyptapis Cockerell, 1909.

Notes-Engel (2001: 54-55) proposed this taxon as subtribe within the Osmiini, but indicated that a future raise of its rank to tribal level was likely and in Engel (2005: 31) and Engel and Perkovsky (2006: 6) it was indeed accepted as tribe as presented in Table 2.

Genus-group names

1839 † Apiaria Germar, Nova Acta Leopoldina, 19: 210. Type species: Apiaria dubia Germar, 1839, by monotypy.

1909 † Ctenoplectrella Cockerell, The Entomologist, 42: 314. Type species: Ctenoplectrella viridiceps Cockerell, 1909, by monotypy.

2001 † Glaesosmia Engel, Bulletin of the American Museum of Natural history, 259: 75. Type species: Glaesosmia genalis Engel, 2001, by original designation.

1909 † Glyptapis Cockerell, The Entomologist, 41: 314. Type species: Glyptapis mirabilis Cockerell, 1909, by monotypy.

2009 † Friccomelissa Wedmann, Wappler & Engel, Naturwissenschaften, 96: 706. Type species: Friccomelissa schopowi Wedman, Wappler & Engel, 2009, by monotypy.

Species-group names

† Ctenoplectrella cockerelli Engel, 2001

2001 † Ctenoplectrella cockerelli Engel, Bulletin of the American Museum of Natural History, 259: 71. Baltic amber inclusion. Type material: Holotype f, B-W 160, AMNH (New York); paratype f, MB.I.1942, ZMHB (Berlin); paratype f, Nr. 1011, Gröhn Collection (Glinde).

† Ctenoplectrella grimaldii Engel, 2001

2001 † Ctenoplectrella grimaldii Engel, Bulletin of the American Museum of Natural History, 259: 73. Baltic amber inclusion. Type material: Holotype f, B-JH 88, AMNH (New York).

† Ctenoplectrella viridiceps Cockerell, 1909

1909 † Ctenoplectrella viridiceps Cockerell, The Entomologist, 42: 314. Baltic amber inclusion. Type material: Neotype f, by designation of Engel (2001: 71), B-JH 95, AMNH (New York).

1931 † Ctenoplectrella dentata Salt, Bernsteinforschungen, 2: 139. Baltic amber inclusion. Type material: Neotype m, by designation of Engel (2001: 71), NB.I.1949, ZMHB (Berlin).-Synonymy in Engel (2001: 69).

1970 † Ctenoplectrella splendens Kelner-Pillault, Annales de la Société Entomologique de France, 6: 13. Baltic amber inclusion. Type material: Holotype m, NB.I.1949, ZMHB (Berlin).-Synonymy in Engel (2001: 69).

† Ctenoplectrella zherikhini Engel & Perkovsky, 2006

2006 † Ctenoplectrella zherikhini Engel & Perkovsky, American Museum Novitates, 3506: 3. Rovno amber inclusion. Type material: Holotype f, UA-1748, UASK (Kiev).

† Friccomelissa schopowi Wedman, Wappler & Engel, 2009

2009 † Friccomelissa schopowi Wedman, Wappler & Engel, Naturwissenschaften, 96: 707. Body fossil from Eocene. Type material: Holotype f, MeI 11036, SMFD (Frankfurt).

† Glaesosmia genalis Engel, 2001

2001 † Glaesosmia genalis Engel, Bulletin of the American Museum of Natural History, 259: 75. Baltic amber inclusion. Type material: Holotype f, B-JH 80, AMNH (New York).

† Glyptapis densopunctata Engel, 2001

2001 † Glyptapis densopunctata Engel, Bulletin of the American Museum of Natural History, 259: 62. Baltic amber inclusion. Type material: Holotype f, B-W 161, AMNH (New York); paratype f, B-JH 89, AMNH (New York); paratype f, Nr. 2623, Gröhn Collection (Glinde); paratype f, Nr. 857, Hoffeins Collection (Hamburg).

† Glyptapis disareolata Engel, 2001

2001 †Glyptapis disareolata Engel, Bulletin of the American Museum of Natural History, 259: 59. Baltic amber inclusion. Type material: Holotype f, B-JH 104, AMNH (New York); paratype f, B-W 156, AMNH (New York).

† Glyptapis fuscula Cockerell, 1909

1909 † Glyptapis fuscula Cockerell, Schriften der physikalisch-ökonomischen Gesellschaft zu Königsberg, 50: 17. Baltic amber inclusion. Type material: Holotype f, K66, GMUG (Göttingen).

1909 † Glyptapis reticulata Cockerell, Schriften der physikalisch-ökonomischen Gesellschaft zu Königsberg, 50: 16. Type material: Neotype f, by designation of Engel (2001: 67), B-JH 78, AMNH (New York).-Synonymy in Engel (2001: 64).

1909 † Glyptapis reducta Cockerell, Schriften der physikalisch-ökonomischen Gesellschaft zu Königsberg, 50: 18. Type material: Holotype f, G4668, GMUG (Göttingen).-Synonymy in Engel (2001: 64).

1931 † Glyptapis neglecta Salt, Bernsteinforschungen, 2: 136. Type material: Neotype f, by designation of Engel (2001: 67), B-JH 78, AMNH (New York).-Synonymy in Engel (2001: 64).

† Glyptapis mirabilis Cockerell, 1909

1909 † Glyptapis mirabilis Cockerell, The Entomologist, 42: 314. Baltic amber inclusion. Type material: Holotype f, K72, GMUG (Göttingen).