{"id":84,"date":"2009-01-21T23:18:28","date_gmt":"2009-01-21T22:18:28","guid":{"rendered":"https:\/\/wpethzprd.ethz.ch\/osmiini\/?page_id=84"},"modified":"2026-05-10T07:12:48","modified_gmt":"2026-05-10T06:12:48","slug":"heriades","status":"publish","type":"page","link":"https:\/\/blogs.ethz.ch\/osmiini\/palaearctic-species\/heriades\/heriades\/","title":{"rendered":"Subgenus Heriades"},"content":{"rendered":"<p style=\"text-align: left\">The subgenus <em>Heriades<\/em> Spinola occurs in the Palaearctic, Afrotropical and Oriental region. It contains 6 described species in the Palaearctic.<\/p>\n<p style=\"text-align: left\"><a href=\"https:\/\/blogs.ethz.ch\/osmiini\/files\/2009\/09\/heriadestruncorum_harz_web1.jpg\"><img loading=\"lazy\" decoding=\"async\" class=\"alignnone size-full wp-image-2568\" src=\"https:\/\/blogs.ethz.ch\/osmiini\/files\/2009\/09\/heriadestruncorum_harz_web1.jpg\" alt=\"\" width=\"433\" height=\"296\" \/><\/a><\/p>\n<h5><span style=\"font-weight: normal\">Nest of <em>Heriades (Heriades) truncorum<\/em> (Linnaeus, 1758) in a preexisting burrow of a pithy stem; the partitions between the brood cells consist of resin. Foto A. Krebs.<\/span><\/h5>\n<h3 style=\"text-align: left\"><strong>Species accounts<\/strong><\/h3>\n<h3 style=\"text-align: left\"><strong><em>Heriades (Heriades) crenulata<\/em><em> <\/em>Nylander, 1856<\/strong><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1856 <em>Heriades <strong>crenulatus<\/strong><\/em> Nylander, Memoires de la Soci\u00e9t\u00e9 des Sciences Naturelles de Cherbourg, 4: 111. <span style=\"text-decoration: underline\">Type material<\/span>: f(f), &#8220;prope Parisios&#8221; [France].<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>A, AZ, BG, BIH, BY,<strong> <\/strong>CH,CZ, D,E, E(Ba), <span style=\"text-decoration: underline\">F<\/span>, F(Co), GE, GR, GR(Aegean Islands, Cr), H, HR, I, I(Sa), I(Si), M, MD, MK, P, PL, RO, RUS(CR,SR), SCG, SK, SLO, UA, <span style=\"font-style: normal\">UA(Crimea)<\/span>; Caucasus<em>. <\/em><strong>Northern Africa: <\/strong>DZ, MA, TN. <strong>Northern Asia: <\/strong>KZ. <strong>Southwestern Asia: <\/strong>KZ, TR.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-Keys, Descriptions: <\/strong>Amiet et al. (2004); Banaszak and Romasenko (2001); Benoist (1929a); Friese (1898); Medvedeva (1978); Scheuchl (1996); Schletterer (1889).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: insect burrows and drilled borings in dead wood; hollow stems (e.g. <em>Phragmites<\/em>) and drilled borings in stems; glass tubes. <strong>Nesting material: <\/strong>Cell partitions and nest plug made of resin. Small pebbles, sand grains, wood fibers and other particles are embedded in the outer plug surface. Cleptoparasite: <em>Stelis breviuscula<\/em>. (Banaszak and Romasenko, 2001; Benoist, 1929a; Brechtel, 1986; Ferton, 1895; Grandi, 1961;  Ruszkowski et al., 1995; Westrich, 1989; Baldock et al., 2008; Ivanov et al., 2019)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Flower preferences: <\/strong>Oligolectic on Asteraceae (Asteroideae, Carduoideae, Cichorioideae) with a preference for the subfamily Asteroideae (Amiet et al., 2004; Westrich, 1989; A. M\u00fcller, unpublished, based on 2 pollen samples from 2 different localities).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Heriades (Heriades) fertoni<\/em><em> <\/em>Benoist, 1938<\/strong><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1938 <em>Heriades <strong>Fertoni<\/strong><\/em> Benoist, Bulletin de la Soci\u00e9t\u00e9 Entomologique de France, 43: 85. <span style=\"text-decoration: underline\">Type material<\/span>: f(f), &#8220;Dellys&#8221; [Algeria], MNHN (Paris).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Africa: <\/strong><span style=\"text-decoration: underline\">DZ<\/span>, TN.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known. <strong>Note: <\/strong>Male unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Flower preferences: <\/strong>Probably oligolectic on Asteraceae (A. M\u00fcller, unpublished, based on 3 pollen samples from 3 different localities).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Heriades (Heriades) labiata <\/em>P\u00e9rez, 1895<\/strong><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1895 <em>Heriades <strong>labiatus<\/strong><\/em> P\u00e9rez, Esp\u00e8ces nouvelles de mellif\u00e8res de Barbarie, p. 21. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Barcelone&#8221; [Spain], MNHN (Paris).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong><span style=\"text-decoration: underline\">E<\/span>. <strong>Northern Africa: <\/strong><del>DZ.<\/del><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known. <strong>Note: <\/strong>Male unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Flower preferences: <\/strong>Unknown.<\/p>\n<h3 style=\"text-align: left\"><strong><em>Heriades (Heriades) rubicola <\/em>P\u00e9rez, 1890<\/strong><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1890 <em>Heriades <strong>rubicola<\/strong><\/em> P\u00e9rez, Actes de la Soci\u00e9t\u00e9 Linn\u00e9enne de Bordeaux, 44: 190. <span style=\"text-decoration: underline\">Type material<\/span>: Syntypes m(m), f(f), &#8220;Bordeaux, Toulouse&#8221; [France], MNHN (Paris).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1892 <em>Heriades<\/em> <strong><em>copetica<\/em><\/strong> Morawitz, Horae Societatis Entomologicae Rossicae (St. Petersburg), 26: 149. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype m, &#8220;Germab&#8221; [Kazakhstan], ZIN (St. Petersburg).-Synonymy in Zanden (1994a: 170).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1931 <em>Heriades<\/em> <em>crenulatus<strong> aurantifer<\/strong><\/em> Cockerell, The Entomologist (London), 64: 159. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype f, by designation of Zanden (1994a: 170), &#8220;Asni&#8221; [Morocco], BMNH (London); paralectotype m.-Synonymy in Zanden (1994a: 170).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1935 <em>Heriades <strong>hierosolomita<\/strong><\/em> Benoist, Bulletin de la Soci\u00e9t\u00e9 Entomologique de France, 40: 277. <span style=\"text-decoration: underline\">Type material<\/span>: f(f), &#8220;Palestine: J\u00e9rusalem&#8221; [Israel], MNHN (Paris).-New synonymy based on type material (A. M\u00fcller, unpublished).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>A, ARM, AZ, B, BG, CH, CZ, D, E, E(Ba), <span style=\"text-decoration: underline\">F<\/span>, F(Co), GB, GR, GR(Aegean Islands, Cr), H, HR, I, I(Sa), I(Si), M, MK, P, RO, RUS(SR), SCG, SK, SLO, UA, <span style=\"font-style: normal\">UA(Crimea)<\/span>. <strong>Northern Africa: <\/strong>DZ, MA, TN. <strong>Northern Asia: <\/strong>KS, KZ, TM<em>. <\/em><strong>Southwestern Asia: <\/strong>CY, IL, IR, IRQ, JOR, RL, SA, SYR, TR.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Amiet et al. (2004); Benoist (1929a); Cross and Notton (2017); Friese (1898); Popov (1960b: 254-255).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: hollow stems (e.g. <em>Phragmites, Rubus<\/em>); old reed galls of <em>Lipara<\/em> flies with 1-7 linearly arranged brood cells; insect burrows in dead wood.<span style=\"font-style: normal\"><strong> <\/strong><\/span><em><span style=\"font-style: normal\"><strong>Nesting material:<\/strong> Cell partitions and nest plug made of resin; sand grains or particles of chewed plant tissue are embedded in the plug and the partitions. <strong>Brood parasites: <\/strong><em>Stelis breviuscula <\/em><\/span><\/em><span style=\"font-style: normal\">(Megachilidae).<\/span><em><span style=\"font-style: normal\"> (Astapenkova et al., 2017; Benoist, 1929a; Bogusch et al., 2015; Cross and Notton, 2017; Ferton, 1894)<\/span><\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Flower preferences: <\/strong>Oligolectic on Asteraceae (Amiet et al., 2004).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Heriades (Heriades) truncorum <\/em>(Linnaeus, 1758)<\/strong><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1758 <em>Apis <strong>truncorum<\/strong><\/em> Linnaeus, Systema Naturae, 10<sup>th<\/sup> edn, vol. 1, p. 575. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype f, by designation of Kirby (1802: 259) [see also Day (1979: 75)], &#8220;in Europa&#8221; [Sweden], LSUK (London). <span style=\"text-decoration: underline\">Type species<\/span> of <em>Heriades<\/em> Spinola.-Combination <em>Hylaeus truncorum<\/em> (Linnaeus) in Fabricius (1793: 305). Combination <em>Megachile truncorum<\/em> (Linnaeus) in Walckenaer (1802: 135). Combination <em>Anthophora truncorum<\/em> (Linnaeus) in Fabricius (1804: 379). Combination <em>Anthidium truncorum<\/em> (Linnaeus) in Panzer (1806: 252). Combination <em>Trachusa truncorum<\/em> (Linnaeus) in Jurine (1807: 252). Combination <em>Trypetes truncorum<\/em> (Linnaeus) in Schenck (1861: 347).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1802 <em>Apis<\/em> <strong><em>xanthogastria<\/em><\/strong> Schrank, Fauna Boica, vol. 2, p. 381. <span style=\"text-decoration: underline\">Type material<\/span>: f(f), &#8220;um Ingolstadt&#8221; [Germany].-Synonymy in Warncke (1986: 115).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1808<strong> <em>Heriades sinuata<\/em> <\/strong>Spinola, Insectorum Liguriae, vol. 2, p. 59. <u>Type material<\/u>: Holotype m, \u201eHabitat prope Genuam\u201c [Italy]. Synonymy in Praz (2017).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1841 <em>Osmia<strong> punctatissima<\/strong><\/em> Lepeletier, Histoire Naturelle des Insectes, Hym\u00e9nopt\u00e8res, vol. 2, p. 329 [not seen]. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype m, by designation of Peters (1971: 51), &#8220;environs de Paris&#8221; [France], MNHN (Paris).-Synonymy in Benoist (1931: 44).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1889 <em>Heriades<\/em> <strong><em>mordax<\/em><\/strong> Schletterer, Zoologisches Jahrbuch f\u00fcr Systematik, 4: 661. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, [North Africa?], ZMHB (Berlin).-Synonymy in Schwarz <em>et al.<\/em> (1996: 115).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>A, AL, AND, ARM, AZ, B, BG, BHI, BY,<strong> <\/strong>CH, CZ, D, DK, E, EST, F, F(Co), FIN, FL, GE, GB, GR, GR(Aegean Islands), H, HR, I, I(Sa), I(Si), L, LT, LV, MD, MK, N, NL, P, PL, RO, RUS(NR,CR,SR), <span style=\"text-decoration: underline\">S<\/span>, SCG, SK, SLO, UA, <span style=\"font-style: normal\">UA(Crimea)<\/span>. <strong>Northern Africa: <\/strong>DZ, MA.<strong> Northern Asia: <\/strong>KS, KZ, RUS(FS), RUS(WS), TJ, UZ<em>. <\/em><strong>Southwestern Asia: <\/strong>CY, IL, IR, RL, SYR, TR.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Amiet et al. (2004); Banaszak and Romasenko (2001); Benoist (1929a); Friese (1898); Medvedeva (1978); Romankova (1995); Scheuchl (1996); Schletterer (1889).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities:insect burrows and drilled borings in dead wood; hollow stems (e.g. <em>Phragmites, Rubus, Vitis<\/em>) and drilled borings in stems; insect burrows in soil (e.g. <em>Odynerus<\/em>); holes in walls; glass tubes. <strong>Nesting material: <\/strong>Cell partitions and nest plug are made of resin (mainly from coniferous trees, also from <em>Prunus avium<\/em>). Small pebbles, sand grains, wood fibers (gnawed by the females from wood) and other particles are usually embedded in the outer plug surface, rarely also in the partitions. Resin is also used to smooth out irregularities or to partially line the cell walls. (Banaszak and Romasenko, 2001; Benoist, 1929a; Bonelli, 1967d; Bosch, Vicens and Blas, 1993; Brechtel, 1986; Enslin, 1933; Ferton, 1894; Friese, 1923; Grandi, 1961; K\u00e4pyl\u00e4, 1978; Le Goff, 2003b; Maciel de Almeida Correia, 1980, 1981b; M\u00fcller et al., 1997; Rozen and Praz, 2016; Ruszkowski et al., 1995; Stoeckhert, 1933; Westrich, 1989; Ivanov et al., 2019; G. Else (unpublished manuscript)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Flower preferences: <\/strong>Oligolectic on Asteraceae with a distinct preference for the subfamily Asteroideae (Amiet et al., 2004; Maciel de Almeida Correia, 1981a; K\u00e4pyl\u00e4, 1978; Westrich, 1989).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Heriades (Heriades) wilmattae <\/em>Cockerell, 1931<\/strong><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1931 <em>Heriades <strong>wilmattae<\/strong><\/em> Cockerell, The Entomologist (London), 64: 159. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype f, by designation of Zanden (1994a: 169), &#8220;Arround&#8221; [Morocco], BMNH (London).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1931 <em>Heriades<\/em> <strong><em>politulus<\/em><\/strong> Cockerell, The Entomologist (London), 64: 160. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype f, by designation of Zanden (1994a: 169), &#8220;Ifrane&#8221; [Morocco], BMNH (London).-Synonymy in Zanden (1994a: 169).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Africa: <\/strong><span style=\"text-decoration: underline\">MA<\/span>.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Flower preferences: <\/strong>Probably oligolectic on Asteraceae (A. M\u00fcller, unpublished, based on 6 pollen samples from 1 locality).<\/p>\n","protected":false},"excerpt":{"rendered":"<p>The subgenus Heriades Spinola occurs in the Palaearctic, Afrotropical and Oriental region. It contains 6 described species in the Palaearctic. Nest of Heriades (Heriades) truncorum (Linnaeus, 1758) in a preexisting burrow of a pithy stem; the partitions between the brood cells consist of resin. Foto A. Krebs. Species accounts Heriades (Heriades) crenulata Nylander, 1856 1856 [&hellip;]<\/p>\n","protected":false},"author":1092,"featured_media":0,"parent":38,"menu_order":0,"comment_status":"open","ping_status":"closed","template":"","meta":{"footnotes":""},"class_list":["post-84","page","type-page","status-publish","hentry"],"_links":{"self":[{"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/pages\/84","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/pages"}],"about":[{"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/types\/page"}],"author":[{"embeddable":true,"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/users\/1092"}],"replies":[{"embeddable":true,"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/comments?post=84"}],"version-history":[{"count":7,"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/pages\/84\/revisions"}],"predecessor-version":[{"id":7439,"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/pages\/84\/revisions\/7439"}],"up":[{"embeddable":true,"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/pages\/38"}],"wp:attachment":[{"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/media?parent=84"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}