{"id":2683,"date":"2009-09-07T13:33:12","date_gmt":"2009-09-07T12:33:12","guid":{"rendered":"https:\/\/wpethzprd.ethz.ch\/osmiini\/?page_id=2683"},"modified":"2026-04-09T15:02:56","modified_gmt":"2026-04-09T14:02:56","slug":"subgenus-osmiaa","status":"publish","type":"page","link":"https:\/\/blogs.ethz.ch\/osmiini\/palaearctic-species\/osmia\/subgenus-osmiaa\/","title":{"rendered":"Subgenus Osmia"},"content":{"rendered":"<p style=\"text-align: left\">The subgenus <em>Osmia<\/em> Panzer is distributed both in the Palaearctic and the Nearctic region. It contains 27 described species, 25 of which occur in the Palaearctic. The North American species <em>Osmia lignaria<\/em> Say, which has been introduced at one locality in Spain (Ortiz-S\u00e1nchez, 2011), is not listed below. Due to the close morphological resemblance, the species of the former subgenera <em>Monosmia<\/em> and <em>Orientosmia<\/em> are included here in the subgenus <em>Osmia<\/em>, which was found to be paraphyletic in its former circumscription (Haider et al., 2013).<\/p>\n<p style=\"text-align: left\"><a href=\"https:\/\/blogs.ethz.ch\/osmiini\/files\/2009\/09\/osmiacerinthidis_web1.jpg\"><img loading=\"lazy\" decoding=\"async\" class=\"alignnone size-full wp-image-2706\" src=\"https:\/\/blogs.ethz.ch\/osmiini\/files\/2009\/09\/osmiacerinthidis_web1.jpg\" alt=\"\" width=\"433\" height=\"296\" \/><\/a><\/p>\n<h5 style=\"text-align: left\"><span style=\"font-weight: normal\">Female of <em>Osmia (Osmia) cerinthidis<\/em> Morawitz, 1875 collecting pollen on <em>Cerinthe <\/em>(Boraginaceae), the species&#8217; exclusive pollen source. Foto A. Gogala.<\/span><\/h5>\n<h3 style=\"text-align: left\">Species accounts<\/h3>\n<h3 style=\"text-align: left\"><span style=\"text-decoration: underline\"><em>apicata<\/em> species group<\/span> (&#8220;<em>Monosmia<\/em>&#8221; and &#8220;<em>Orientosmia<\/em>&#8220;)<\/h3>\n<h3><strong><em>Osmia (Monosmia) apicata<\/em><\/strong><strong> <\/strong><span style=\"font-weight: normal\">Smith, 1853<\/span><\/h3>\n<p style=\"text-align: left;padding-left: 30px\">1853 <em>Osmia <strong>apicata<\/strong><\/em> Smith, Catalogue of Hymenopterous Insects in the Collection of the British Museum, vol. 1, p. 140. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype f, by designation of Tkalcu (1974b: 339), &#8220;Albania&#8221; [Albania], BMNH (London). <span style=\"text-decoration: underline\">Type species<\/span> of <em>Monosmia<\/em> Tkalcu.<\/p>\n<p style=\"text-align: left;padding-left: 30px\">1869 <em>Osmia<\/em> <strong><em>macroglossa<\/em><\/strong> Gerstaecker, Entomologische Zeitung (Stettin), 30: 349. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype f, by designation of Tkalcu (1974b: 339), &#8220;Griechenland&#8221; [Greece], ZMHB (Berlin).-Combination <em>Osmia (Aceratosmia) macroglossa<\/em> Gerstaecker in Schmiedeknecht (1885: 19 [885]). Combination <em>Osmia (Pachyosmia) macroglossa<\/em> Gerstaecker in Friese (1911b: 124).-Synonymy in Tkalcu (1974b: 339).<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Distribution-Europe:<\/strong><em> <\/em><span style=\"text-decoration: underline\">AL<\/span>, ARM, AZ, BG, BIH, I, GE, GR, GR(Aegean Islands), HR, MK, RUS(SR), SCG, SLO.<em> <\/em><strong>Southwestern Asia:<\/strong><em> <\/em>IL, IR, IRQ, JOR, SYR, TR.<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Identification-<\/strong><strong>Keys, Descriptions:<\/strong><strong> <\/strong>Ducke (1900);<strong> <\/strong>Medvedeva (1978); Peters (1978a), Schmiedeknecht (1885-1886); Tkalcu (1974b: 337-339).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: crevices in stone walls. <strong>Nesting material: <\/strong>On the Peloponnese in Greece in 2024, numerous females were observed to enter holes in a vertical earth bank, where they collected moist soil, indicating that mud is used as nest-building material; hundreds of females of <em>Osmia bicornis<\/em> and <em>O. cornuta<\/em> exploited the same mining site for mud extraction (T. Wood, personal communication). (Ducke, 1900; Graeffe, 1902)<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Flower preferences:<\/strong> Oligolectic on Boraginaceae with a strong preference for <em>Onosma<\/em> (Teppner, 1996; Gogala and Surina, 2011; Haider et al., 2013).<\/p>\n<h3><strong><em>Osmia (Orientosmia) maxillaris<\/em><\/strong><strong> <\/strong><span style=\"font-weight: normal\">Morawitz<\/span><span style=\"font-weight: normal\">, 1875<\/span><\/h3>\n<p style=\"text-align: left;padding-left: 30px\">1875 <em>Osmia <strong>maxillaris<\/strong><\/em> Morawitz, Travel to Turkestan by A. P. Fedtschenko, p. 82. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype m, by designation of Peters (1978a: 317) &#8220;in valle Sarafschan, Warsaminor, Obburden, Iskander&#8221; [Tajikistan], ZIN (St. Petersburg); paralectotypes m, ff. <span style=\"text-decoration: underline\">Type species<\/span> of <em>Orientosmia<\/em> Peters.-Combination <em>Osmia (Pachyosmia) maxillaris<\/em> Morawitz in Friese (1911b: 125).<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Distribution-Northern Asia:<\/strong><em> <\/em>KS, KZ, <span style=\"text-decoration: underline\">TJ<\/span>, UZ<em>.<\/em><\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Identification-<\/strong><strong>Keys, Descriptions:<\/strong><strong> <\/strong>Ducke (1900);<strong> <\/strong>Peters (1978a); M\u00fcller (2012b).<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Nesting biology: <\/strong>Unknown.<strong><br \/>\n<\/strong><\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Flower preferences: <\/strong>Oligolectic on Fabaceae with long flower corollas, e.g. <em>Astragalus<\/em> (M\u00fcller, 2012b; Haider et al., 2013).<\/p>\n<h3 style=\"text-align: left\"><em>Osmia (Orientosmia) <\/em><strong><em>maxschwarzi<\/em> <\/strong><span style=\"font-weight: normal\">M\u00fcller, 2012<\/span><\/h3>\n<p style=\"text-align: left;padding-left: 30px\">2012 <em>Osmia (Orientosmia) <\/em><strong><em>maxschwarzi<\/em><\/strong> M\u00fcller, Journal of the Swiss Entomological Society, 85: 29. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype m, \u201eFiruzkuh county, Markazi district, 28km E of Firuzkuh, Peynap valley, 2450m\u201c [Iran], ETHZ (Z\u00fcrich); paratypes mm, ff.<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Distribution-Southwestern Asia:<\/strong><em> <\/em><span style=\"text-decoration: underline\">IR<\/span>, IRQ, TR.<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Identification-Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known.<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Flower preferences: <\/strong>Oligolectic on Fabaceae with long flower corollas, e.g. <em>Astragalus<\/em> (M\u00fcller, 2012b; Haider et al., 2013).<\/p>\n<h3><em>Osmia (Orientosmia) <\/em><strong><em>scheherazade<\/em> <\/strong><span style=\"font-weight: normal\">Peters, 1978<\/span><\/h3>\n<p style=\"text-align: left;padding-left: 30px\">1978 <em>Osmia (Orientosmia) maxillaris <\/em><strong><em>scheherazade<\/em><\/strong> Peters, Senckenbergiana Biologica (Frankfurt), 58: 319. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype m, \u201eG\u00fcr\u00fcn\u201c [Turkey], SMFD (Frankfurt); paratypes mm, ff.<\/p>\n<p style=\"text-align: left;padding-left: 30px\">1978 <em>Osmia (Orientosmia) maxillaris <\/em><strong><em>dinazade<\/em><\/strong> Peters, Senckenbergiana Biologica (Frankfurt), 58: 319. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype m, \u201ePul-i-chatum (an der Grenze von Turkmenien)\u201c [Iran], ZIN (St. Petersburg); paratype m.-Synonymy in M\u00fcller (2012b: 28).<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Distribution-Europe:<\/strong> AZ, RUS(SR). <strong>Southwestern Asia:<\/strong> IR, <span style=\"text-decoration: underline\">TR<\/span>.<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Identification-Keys, Descriptions: <\/strong>M\u00fcller (2012b).<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Flower preferences: <\/strong>Possibly oligolectic on Fabaceae with long flower corollas, e.g. <em>Astragalus<\/em> (M\u00fcller, 2012b; Haider et al., 2013).<\/p>\n<h3 style=\"text-align: left\"><span style=\"text-decoration: underline\"><strong><em>bicornis<\/em> species group<\/strong><\/span><\/h3>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Osmia) ariadne <\/em><\/strong><span style=\"font-weight: normal\">Peters, 1978<\/span><\/h3>\n<p style=\"text-align: left;padding-left: 30px\">1978 <em>Osmia (Osmia) <strong>ariadne<\/strong><\/em> Peters, Senckenbergiana Biologica (Frankfurt), 58: 296. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;W-Kreta, Meskla&#8221; [Greece: Crete], SMFD (Frankfurt); paratype f.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong><span style=\"text-decoration: underline\">GR(Cr)<\/span><em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Zanden (1986: 72).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<strong><br \/>\n<\/strong><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Unknown.<\/p>\n<h3 style=\"text-align: left\"><strong><em><span style=\"font-style: normal;font-weight: normal\"><strong><em>Osmia (Osmia) bicornis <\/em><\/strong>(Linnaeus, 1758)<\/span><\/em><\/strong><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1758 <em>Apis<\/em> <strong><em>bicornis<\/em><\/strong> Linnaeus, Systema Naturae, 10th edn, vol. 1, p. 575. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;in Europa&#8221; [Sweden], LSUK (London). <span style=\"text-decoration: underline\">Type species<\/span> of <em>Osmia<\/em> Panzer.-Combination <em>Osmia (Ceratosmia) bicornis<\/em> (Linnaeus) in Schmiedeknecht (1885: 19 [885]).-Synonymy with <em>Osmia rufa<\/em> (Linnaeus) in Dalla Torre (1896: 407) and in Warncke (1986: 121), but see Day (1979: 51, 70).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1758 <em>Apis <strong>rufa<\/strong><\/em> Linnaeus, Systema Naturae, 10<sup>th<\/sup> edn, vol. 1, p. 575. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype m, by designation of Day (1979: 70), &#8220;in Europa&#8221; [Sweden], LSUK (London).-Combination <em>Osmia (Pachyosmia) rufa<\/em> (Linnaeus) in Friese (1911b: 126). Combination <em>Osmia (Osmia) rufa <\/em>(Linnaeus) in Zanden (1988b: 127).-Synonymy in Kirby (1802: 273) and in Day (1979: 51, 70).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>A, AL, AND, ARM, AZ, B, BG, BIH, BY,<strong> <\/strong>CH, CZ, D, DK, E, E(Ba), EST, F, F(Co), FIN, FL, GB, GE, GR, GR(Aegean Islands), H, HR, I, I(Sa), I(Si), IRL, L, LT, LV, MK, N, NL, P, PL, RO, RUS(NR,CR,SR), <span style=\"text-decoration: underline\">S<\/span>, SCG, SK, SLO, UA, UA(Crimea)<em>. <\/em><strong>Northern Africa: <\/strong>DZ, MA, TN.<strong> Northern Asia: <\/strong>KS, KZ, RUS(FS), TM; <span style=\"text-decoration: line-through\">J<\/span><em>. <\/em><strong>Southwestern Asia: <\/strong>CY, IL, IR, RL, SYR, TR<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Amiet et al. (2004); Banaszak and Romasenko (2001); Benoist (1931); Ducke (1900); Medvedeva (1978); M\u00f3cz\u00e1r (1958); Peters (1978a); Radoszkowski (1887a: 279); Romankova (1984); Scheuchl (1996); Schmiedeknecht (1885-1886).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities of most diverse form and size: insect burrows in dead wood and drilled borings in wooden blocks; hollow stems (e.g. <em>Arundo, Phragmites<\/em>) and drilled borings in pithy stems; abandoned cells in exposed nests of other aculeates (e.g. <em>Chalicodoma<\/em>); insect burrows in horizontal or vertical ground (e.g. of <em>Anthophora, Colletes<\/em>); crevices in walls; cracks between shingles on house walls; key holes, pocket watches and flutes; glass tubes; empty snail shells. <strong>Nesting material: <\/strong>Cell partitions and nest plug are made of mud. In larger cavities, the cells consist entirely or partially of mud. In rare cases, nests inside cavities are entirely composed of mud, then similar to the nests of <em>Chalicodoma. <\/em>Irregularities inside the nests are often smoothed with mud. (Banaszak and Romasenko, 2001; Benoist, 1931; Bonelli, 1966, 1967a, 1968; Brechtel, 1986; Friese, 1923; Grandi, 1961; Ivanov, 2006; Le Goff, 2006; Malyshev, 1937; M\u00fcller et al., 1997; Olifir, 1980; O&#8217;Toole and Raw, 1999; Raw, 1972; Schmiedeknecht, 1885-1886; Smith, 1844; Tasei, 1973; Vicens, Bosch and Blas, 1993; Westrich, 1989; Ivanov et al., 2019; G. Else, unpublished manuscript)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Polylectic; pollen sources include plants belonging to 19 different families; flowers containing no or very little nectar, such as <em>Ranunculus<\/em> (Ranunculaceae), <em>Quercus<\/em> (Fabaceae), <em>Papaver<\/em> (Papaveraceae) or Cistaceae, are the predominant pollen hosts (Amiet et al., 2004; Westrich, 1989; Haider et al., 2013).<\/p>\n<h3 style=\"text-align: left\"><em> <span style=\"font-style: normal\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>bicornis <\/strong><\/em><span style=\"font-weight: normal\">(Linnaeus, 1758)<\/span><\/span><\/em><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1758 Automatically established nominotypical subspecific taxon (original description and type specimen are identical with those of the species rank taxon of the same name above).-Combination <em>Osmia rufa rufa<\/em> (Linnaeus) in Peters (1978a: 290).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1776<em> Apis <strong>agino<\/strong><\/em> Harris, Exposition of English Insects, p. 163. <span style=\"text-decoration: underline\">Type material<\/span>: m(m), [England].-Synonymy in Peters (1978a: 290).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1844 <em>Osmia <strong>hedera<\/strong><\/em> Smith, The Zoologist (London), 2: 747. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype m, &#8220;Hampshire&#8221; [United Kingdom].-Synonymy in Tkalcu (1977b: 232).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1899 <em>Osmia<\/em> <em>rufa<strong> borealis<\/strong><\/em> Ducke, Entomologische Nachrichten (Berlin), 25: 215. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;aus Norwegen&#8221; [Norway], NMW (Wien).-Combination <em>Osmia (Pachyosmia) rufa<\/em> <em>borealis<\/em> Ducke in Friese (1911b: 126).-Synonymy in Tkalcu (1977b: 232).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe:<\/strong><em> <\/em>AND, DK, E, F(Co), FIN, GB, I(Sa), I(Si), IRL, N, P, <span style=\"text-decoration: underline\">S<\/span><em>. <\/em><strong>Southwestern Asia:<\/strong> IR.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Ducke (1900); Peters (1978a).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>globosa <\/strong><\/em><span style=\"font-weight: normal\">(Scopoli, 1763)<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1763 <em>Apis <strong>globosa<\/strong><\/em> Scopoli, 1763, Entomologia Carniolica, p. 300. <span style=\"text-decoration: underline\">Type material<\/span>: Neotype m, by designation of Warncke (1986: 74), &#8220;Klagenfurt, K\u00e4rnten&#8221; [Austria], ZSM (M\u00fcnchen).-Combination <em>Osmia rufa globosa<\/em> (Scopoli) in Warncke (1986: 100).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1790 <em>Apis <strong>cornigera<\/strong><\/em> Rossi, Fauna Etrusca, vol. 2, p. 108. <span style=\"text-decoration: underline\">Type material<\/span>: f(f), [Italy].-Combination <em>Osmia rufa cornigera<\/em> (Rossi) in Peters (1978a: 294).-Synonymy in Warncke (1986: 94).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1799 <em>Apis<strong> fronticornis<\/strong><\/em> Panzer, Faunae Insectorum Germanicae, 63: 20. <span style=\"text-decoration: underline\">Type material<\/span>: f(f), [Germany].-Synonymy with <em>Osmia rufa cornigera<\/em> (Rossi) in Peters (1978a: 294). Synonymy with <em>Osmia rufa globosa<\/em> (Scopoli) in Warncke (1986: 98).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe:<\/strong> A, AL, ARM, AZ, B, BG, BIH, BY,<strong> <\/strong>CH, CZ, D, DK, E, EST, F, F(Co), FL, GE, GR, GR(Aegean Islands), H, HR, <span style=\"text-decoration: underline\">I<\/span>, I(Si), L, LT, LV, MK, NL, PL, RO, RUS(NR,CR,SR), S, SCG, SK, SLO, UA, UA(Crimea). <strong>Northern Africa: <\/strong>TN.<strong> Northern Asia: <\/strong>KS, KZ, RUS(FS), TM. <strong>Southwestern Asia:<\/strong> CY?, IL, IR, RL, SYR, TR.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Peters (1978a); Radoszkowski (1887a: 280).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>fracticornis <\/strong><\/em><span style=\"font-weight: normal\">P\u00e9rez, 1895<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1895 <em>Osmia <strong>fracticornis<\/strong><\/em> P\u00e9rez, Esp\u00e8ces nouvelles de mellif\u00e8res de Barbarie, p. 10. <span style=\"text-decoration: underline\">Type material<\/span>: Syntypes m(m), f(f), [Algeria], MNHN (Paris).-Combination <em>Osmia (Pachyosmia) fracticornis<\/em> P\u00e9rez in Friese (1911b: 126). Combination <em>Osmia rufa fracticornis<\/em> P\u00e9rez in Peters (1978a: 295).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe:<\/strong> E(Ba), I(Sa), I(Si). <strong>Northern Africa: <\/strong><span style=\"text-decoration: underline\">DZ<\/span>, MA, TN<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Ducke (1900); Peters (1978a).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Osmia) cerinthidis <\/em><\/strong><span style=\"font-weight: normal\">Morawitz, 1875<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1875 <em>Osmia <strong>cerinthidis<\/strong><\/em> Morawitz, Horae Societatis Entomologicae Rossicae (St. Petersburg), 12: 41. <span style=\"text-decoration: underline\">Type material<\/span>: f(f), &#8220;Borshom&#8221; [Georgia], ZIN (St. Petersburg).-Combination <em>Osmia (Ceratosmia) cerinthidis<\/em> Morawitz in Schmiedeknecht (1885: 19 [885]). Combination <em>Osmia (Pachyosmia) rufa<\/em> <em>cerinthidis<\/em> Morawitz in Friese (1911b: 126).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>A, ARM, AZ, BG, CZ, D, F, <span style=\"text-decoration: underline\">GE<\/span>, GR, H, HR, I, I(Si), MK, PL, RO, RUS(SR), SK, SLO, UA, UA(Crimea); <span style=\"text-decoration: line-through\">B<\/span><em>. <\/em><strong>Northern Asia: <\/strong>RC?<em>. <\/em><strong>Southwestern Asia: <\/strong>IR, TR<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Amiet et al. (2004);Banaszak and Romasenko (2001); Ducke (1900, as <em>Osmia rufa var. cerinthidis<\/em>); Medvedeva (1978); Peters (1978a); Scheuchl (1996); Schmiedeknecht (1885-1886); Wu (2006).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: drilled borings in wooden blocks; hollow stems (e.g. <em>Phragmites<\/em>); abandoned brood cells of <em>Anthophora <\/em>in vertical cliffs. <strong>Nesting material: <\/strong>Cell partitions and nest plug are made of mud. (Banaszak and Romasenko, 2001; Olifir, 1980; Rozen and \u00f4zbek, 2005; Ivanov et al., 2019)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Oligolectic on Boraginaceae with a strong preference for <em>Cerinthe<\/em> (Boraginaceae) (Amiet et al., 2004; Westrich, 1989; Haider et al., 2013).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>cerinthidis <\/strong><\/em><span style=\"font-weight: normal\">Morawitz, 1875<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1875 Automatically established nominotypical subspecific taxon (original description and type specimen are identical with those of the species rank taxon of the same name above).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>A, ARM, AZ, CZ, D, F, <span style=\"text-decoration: underline\">GE<\/span>, GR, H, HR, I, MK, PL, RUS(SR), SK, SLO, UA, UA(Crimea).<strong>Northern Asia: <\/strong>RC<em>. <\/em><strong>Southwestern Asia: <\/strong>IR, TR<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Peters (1978a).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>crassiclypeata <\/strong><\/em><span style=\"font-weight: normal\">Peters, 1978<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1978 <em>Osmia (Osmia) cerinthidis <strong>crassiclypeata<\/strong><\/em> Peters, Senckenbergiana Biologica (Frankfurt), 58: 299. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Palermo&#8221; [Italy: Sicily], Schwarz Collection (Ansfelden).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong><span style=\"text-decoration: underline\">I(Si)<\/span><em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known.<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Osmia) cornifrons <\/em><\/strong><span style=\"font-weight: normal\">(Radoszkowski, 1887)<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1887 <em>Chalicodoma<strong> cornifrons<\/strong><\/em> Radoszkowski, Horae Societatis Entomologicae Rossicae (St. Petersburg), 21: 430. <span style=\"text-decoration: underline\">Type material<\/span>: f(f), &#8220;Kor\u00e9e&#8221; [Korea].-Combination <em>Osmia (Ceratosmia) cornifrons<\/em> (Radoszkowski) in Gussakovskij (1932: 61).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Asia: <\/strong>J, RC, ROK, RUS(FS); <span style=\"text-decoration: underline\">Korea<\/span><em>. <\/em><strong>Extralimital: <\/strong>Nearctic (USA) [Introduced].<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Romankova (1984, 1985b: 943-944, 1995); Wu (2006); Yasumatsu and Hirashima (1950).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: insect borings in dead wood; hollow stems (e.g. <em>Conium, Phragmites<\/em>, bamboo) and straw tubes; space between a folded newspaper. <strong>Nesting material: <\/strong>Cell partitions and nest plug are made of mud. In large cavities, cells are irregularly arranged with cell walls partially or entirely built of mud. (Hirashima, 1973; Kitamura and Maeta, 1969; Maeta, 1969, 1978; Rust, 1974; Yamada et al., 1971)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Polylectic; pollen sources include plants belonging to 9 different families; flowers of Fabaceae are important pollen hosts (Quest, 2009; Haider et al., 2013).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Osmia) cornuta <\/em><\/strong><span style=\"font-weight: normal\">(Latreille, 1805)<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1805 <em>Megachile <strong>cornuta<\/strong><\/em> Latreille, Histoire Naturelle des Crustac\u00e9s et des Insectes, vol. 14: 59 [not seen]. <span style=\"text-decoration: underline\">Type material<\/span>: [France]. Combination <em>Osmia (Ceratosmia) cornuta<\/em> (Latreille) in Schmiedeknecht (1885: 19 [885]). Combination <em>Osmia (Pachyosmia) cornuta<\/em> (Latreille) in Friese (1911b: 127).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe:<\/strong> A, AND, ARM, AZ, B, BG, BIH, BY, CH, CZ, D, DK, E, E(Ba), <span style=\"text-decoration: underline\">F<\/span>, F(Co), FL, GB, GE, GR, GR(Aegean Islands, Cr), H, HR, I, I(Sa), I(Si), L, MK, NL, P, PL, RO, RUS(SR), S, SCG, SK, SLO, UA, UA(Crimea). <strong>Northern Africa:<\/strong> ET, DZ, TN. <strong>Northern Asia:<\/strong> KZ, TM; Turkestan. <strong>Southwestern Asia:<\/strong> CY, IL, IR, RL, SYR, TR.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Amiet et al. (2004); Banaszak and Romasenko (2001); Benoist (1931); Ducke (1900); Medvedeva (1978); M\u00f3cz\u00e1r (1958); Peters (1978a); Radoszkowski (1887a: 291-292); Scheuchl (1996); Schmiedeknecht (1885-1886).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: drilled borings in wooden blocks; hollow stems (e.g. <em>Phragmites<\/em>, bamboo) and drilled borings in pithy stems; crevices and holes in walls, rocks or stones; insect burrows in the soil (e.g. of <em>Anthophora<\/em>); glass tubes; empty snail shells (e.g. of <em>Zonites algirus<\/em>). <strong>Nesting material:<\/strong> Cell partitions and nest plug are made of mud. In larger cavities, cells are partially or entirely made of mud. (Benoist, 1931; Brechtel, 1986; Friese, 1923; Gogala, 1999; Graeffe, 1902; Mavromoustakis, 1948c; M\u00fcller et al., 1997; Olifir, 1980; Popovici-Baznosanu, 1909; Stoeckhert, 1933; Tasei, 1973; Vicens, Bosch and Blas, 1993; Westrich, 1989; Ivanov et al., 2019; G. Le Goff, personal obervation)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Flower preferences: <\/strong>Polylectic; pollen sources include plants belonging to 13 different families; flowers of Rosaceae are the most important pollen hosts (Amiet et al., 2004; Westrich, 1989; Haider et al., 2013).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies<\/span><em> <strong>cornuta <\/strong><\/em><span style=\"font-weight: normal\">(Latreille, 1805)<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1805 Automatically established nominotypical subspecific taxon (original description and type specimen are identical with those of the species rank taxon of the same name above).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe:<\/strong> A, AND, B, BG, BIH, BY, CH, CZ, D, DK, E, E(Ba), <span style=\"text-decoration: underline\">F<\/span>, F(Co), FL, GR, H, HR, I, I(Sa), I(Si), L, MK, NL, P, PL, RO, RUS(SR), S, SCG, SK, SLO, UA, UA(Crimea). <strong>Northern Africa:<\/strong> DZ, TN. <strong>Northern Asia:<\/strong> KZ.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Peters (1978a).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>divergens <\/strong><\/em><span style=\"font-weight: normal\">Friese, 1920<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1920 <em>Osmia <strong>divergens<\/strong><\/em> Friese, Deutsche Entomologische Zeitschrift (Berlin), 1920: 51. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype f, by designation of Peters (1978a: 304), &#8220;Turkestan&#8221; [Turkestan], ZMHB (Berlin).-Combination <em>Osmia (Osmia) cornuta<\/em> <em>divergens<\/em> Friese in Peters (1978a: 304).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Asia: <\/strong>TM; <span style=\"text-decoration: underline\">Turkestan<\/span><em>. <\/em><strong>Southwestern Asia: <\/strong>IR<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Peters (1978a).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies<\/span><em> <strong>neoregaena <\/strong><\/em><span style=\"font-weight: normal\">Mavromoustakis, 1938<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1938 <em>Osmia <strong>neoregaena<\/strong><\/em> Mavromoustakis, The Annals and Magazine of Natural History (London), ser. 11, 2: 471. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Saittas, 2500 ft&#8221; [Cyprus], DAAN (Nicosia); paratypes mm, ff.-Combination <em>Osmia (Osmia) cornuta neoregaena<\/em> Mavromoustakis in Peters (1978a: 304).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution- Europe: <\/strong>GR(Lesbos). <strong>Southwestern Asia:<\/strong> <span style=\"text-decoration: underline\">CY<\/span><em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Peters (1978a).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>quasirufa <\/strong><\/em><span style=\"font-weight: normal\">Peters, 1978<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1978 <em>Osmia (Osmia) cornuta <strong>quasirufa<\/strong><\/em> Peters, Senckenbergiana Biologica (Frankfurt), 58: 303. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Kingi (Maryut, sw von Alexandria) im Garten&#8221; [Egypt], SMFD (Frankfurt); paratypes mm, ff.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>ARM, AZ, GE, GR(Chios, Cr?)<em>. <\/em><strong>Northern Africa: <\/strong><span style=\"text-decoration: underline\">ET<\/span><em>. <\/em><strong>Southwestern Asia: <\/strong>IL, IR, RL, SYR, TR<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known.<\/p>\n<h3><strong><em>Osmia (Osmia) cyrenaica <\/em><\/strong><span style=\"font-weight: normal\">Peters, 1978<\/span><\/h3>\n<p style=\"text-align: left;padding-left: 30px\">1978 <em>Osmia (Osmia) <strong>cyrenaica<\/strong><\/em> Peters, Senckenbergiana Biologica (Frankfurt), 58: 308. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype m, &#8220;Cyrenaica, Cyrene, 1800 feet&#8221; [Libya], BMNH (London); paratype m.<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Distribution-Northern Africa:<\/strong><em> <\/em><span style=\"text-decoration: underline\">LAR<\/span><em>.<\/em><\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Identification-<\/strong><strong>Keys, Descriptions:<\/strong><strong> <\/strong>No supplementary or more detailed morphological description known. <strong>Note: <\/strong>Female unknown.<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Flower preferences: <\/strong>Unknown.<\/p>\n<h3><strong><em>Osmia (Osmia) excavata<\/em><\/strong><strong> <\/strong><span style=\"font-weight: normal\">Alfken, 1903<\/span><\/h3>\n<p style=\"text-align: left;padding-left: 30px\">1903 <em>Osmia (Chalcosmia)<strong> excavata<\/strong><\/em> Alfken, Zeitschrift f\u00fcr Systematische Hymenopterologie und Dipterologie, 3: 210. <span style=\"text-decoration: underline\">Type material<\/span>: f(f), &#8220;Tokio&#8221; [Japan], ZMHB (Berlin).<\/p>\n<p style=\"text-align: left;padding-left: 30px\">1911 <em>Osmia<\/em> <strong><em>mitsukurii<\/em><\/strong> Cockerell, Proceedings of the United States National Museum, 39: 639. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, [Japan], USNM (Washington).-Synonymy in Yasumatsu and Hirashima (1950: 3).<\/p>\n<p style=\"text-align: left;padding-left: 30px\">1931 <em>Osmia (Pachyosmia)<\/em> <strong><em>heudei<\/em><\/strong> Cockerell, American Museum Novitates, 466: 2. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Zi-ka-wei&#8221; [China], AMNH (New York).-Synonymy in Yasumatsu and Hirashima (1950: 3).<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Distribution-Northern Asia:<\/strong><em> <\/em><span style=\"text-decoration: underline\">J<\/span>, RC, ROK.<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Identification-<\/strong><strong>Keys, Descriptions:<\/strong><strong> <\/strong>Wu (2006); Yasumatsu and Hirashima (1950).<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: insect borings in dead wood; hollow stems (e.g. <em>Phragmites<\/em>, bamboo) and straw tubes. <strong>Nesting material: <\/strong>Cell partitions and nest plug are made of mud. (Hirashima, 1957, 1958, 1973; Kitamura and Maeta, 1969; Maeta, 1978)<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Flower preferences: <\/strong>Polylectic; pollen sources include plants belonging to 7 different families; flowers of Brassicaceae are the most important pollen hosts (Haider et al., 2013).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Osmia) kohl<\/em><\/strong><strong><em>ii <\/em><\/strong><span style=\"font-weight: normal\">Ducke, 1899<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1899 <em>Osmia <strong>kohlii<\/strong><\/em> Ducke, Entomologische Nachrichten (Berlin), 25: 213. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype f, by designation of Peters (1978a: 306), &#8220;Sicilien&#8221; [Italy: Sicily], NMW (Wien); paralectotypes mm, ff.-Combination <em>Osmia (Pachyosmia) kohli<\/em> Ducke in Friese (1911b: 127).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong><span style=\"text-decoration: underline\">I(Si<\/span> and adjacent Calabrian mainland), M; <span style=\"text-decoration: line-through\">F(Co)<\/span><em>. <\/em><strong>Northern Africa: <\/strong>DZ?<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Benoist (1931); Ducke (1900); Peters (1978a).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Polylectic; pollen sources include plants belonging to 11 different families; flowers containing no or very little nectar, such as  <em>Papaver<\/em> (Papaveraceae), <em>Ranunculus<\/em> (Ranunculaceae) or Cistaceae, are important pollen hosts (Haider et al., 2013).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Osmia) longicornis <\/em><\/strong><span style=\"font-weight: normal\">Morawitz, 1875<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1875 <em>Osmia <strong>longicornis<\/strong><\/em> Morawitz, Travel to Turkestan by A. P. Fedtschenko, p. 78. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype f, by designation of Tkalcu (1969a: 343), &#8220;Taschkent&#8221; [Uzbekistan], ZMHB (Berlin); paralectotype m.-Combination <em>Osmia (Pachyosmia) longicornis<\/em> Morawitz in Friese (1911b: 125).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution- Europe: <\/strong>Caucasus. <strong>Northern Asia: <\/strong>TJ, <span style=\"text-decoration: underline\">UZ<\/span><em>. <\/em><strong>Southwestern Asia: <\/strong>AFG, PAK<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Ducke (1900); Peters (1978a); Romankova (1985b: 943-944); Tkalcu (1969a: 343).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Probably polylectic; the few available pollen samples contained pollen of Rosaceae and <em>Salix<\/em> (Salicaceae) (Haider et al., 2013).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Osmia) opima <\/em><\/strong><span style=\"font-weight: normal\">Romankova, 1985<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1985 <em>Osmia (Osmia) <strong>opima<\/strong><\/em> Romankova, Zoologicheskiy Zhurnal (Moscow), 64: 942. <span style=\"text-decoration: underline\">Type material<\/span>: Syntypes m(m), f(f), &#8220;Primorye, 14 km north of the village Kievka&#8221; [Russia]<\/p>\n<p style=\"padding-left: 30px;text-align: left\">2004 <em>Osmia (Osmia) <strong>yanbianense<\/strong><\/em> Wu, Acta Zootaxonomica Sinica, 29: 532. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype m, &#8220;Jilin, Yanbian (42.9\u252c\u2591N, 129.5\u252c\u2591E)&#8221; [China], IZCAS (Beijing); paratype f.-New synonymy based on Branstetter et al. (2021) and type material.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Asia: <\/strong>RC(NE), RC(NO), ROK, <u>RUS<\/u>(FS).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Romankova (1995); Wu (2006).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-<\/strong><strong>Nesting site: <\/strong>Preexisting cavities: insect burrows in dead wood. <strong>Nesting material: <\/strong>Cell partitions and nest plug are made of mud. (Romankova, 1985b)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Polylectic (T. Romankova, personal communication).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Osmia) pedicornis <\/em><\/strong><span style=\"font-weight: normal\">Cockerell, 1919<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1919 <em>Osmia (Pachyosmia)<strong> pedicornis<\/strong><\/em> Cockerell, Proceedings of the United States National Museum (Washinton), 55: 182. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Soochow&#8221; [China], USNM (Washington); paratype m.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Asia: <\/strong>J, <u>RC<\/u>(CE), ROK, RUS(FS).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Romankova (1994: 123, 1995); Wu (2006); Yasumatsu and Hirashima (1950).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: insect burrows in dead wood; hollow stems (e.g. <em>Phragmites<\/em>, bamboo) and straw tubes. <strong>Nesting material: <\/strong>Cell partitions and nest plug are made of mud. (Hirashima, 1958; Kitamura and Maeta, 1969; Maeta, 1969, 1978)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Polylectic; pollen sources include plants belonging to 7 different families; flowers containing no or very little nectar, such as <em>Quercus<\/em> (Fabaceae), <em>Ranunculus<\/em> (Ranunculaceae) or <em>Juglans<\/em> (Juglandaceae), are important pollen hosts  (Quest, 2009; Haider et al., 2013).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Osmia) <\/em><\/strong><strong><em>quadricornuta<\/em><\/strong> <span style=\"font-weight: normal\">Wu, 2004<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">2004 <em>Osmia <strong>quadricornuta<\/strong><\/em> Wu, Acta Zootaxonomica Sinica, 29: 531. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Yunnan, Kunming Xishan (25\u252c\u2591N, 102.2\u252c\u2591E), 2100 m&#8221; [China], IZCAS (Beijing); paratype m.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Asia: <\/strong><span style=\"text-decoration: underline\">RC(SW)<\/span>.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Wu (2006).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Unknown.<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Osmia) rufina<\/em><\/strong><span style=\"font-weight: normal\"> Cockerell, 1931<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1931 <em>Osmia (Pachyosmia) <strong>rufina<\/strong><\/em> Cockerell, American Museum Novitates (New York), 466: 3. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Z\u00f2-S\u00e8&#8221; [China], AMNH (New York).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">2004 <em>Osmia <strong>rufinoides<\/strong><\/em> Wu, Acta Zootaxonomica Sinica, 29: 532. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Shanghai&#8221; [China], IZCAS (Beijing); paratypes ff.-New synonymy based on Branstetter et al. (2021).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Asia:<\/strong><span style=\"text-decoration: underline\">RC<\/span><em>, <\/em>RC(CE)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Wu (2006).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Unknown.<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Osmia) taurus <\/em><\/strong><span style=\"font-weight: normal\">Smith, 1873<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1873 <em>Osmia <strong>Taurus<\/strong><\/em> Smith, Transactions of the Entomological Society of London, 1873: 201. <span style=\"text-decoration: underline\">Type material<\/span>: Syntypes m(m), f(f),&#8221;Hiogo&#8221; [Japan], BMNH (London).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1911 <em>Osmia<\/em> <strong><em>matsumurae<\/em><\/strong> Cockerell, Proceedings of the United States National Museum, 39: 640. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype m, &#8220;Sapporo&#8221; [Japan], USNM (Washington).-Synonymy in Yasumatsu and Hirashima (1950: 2).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Asia: <\/strong><u>J<\/u>, RC, ROK, RUS(FS)<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Romankova (1984, 1995); Wu (2006); Yasumatsu and Hirashima (1950).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: insect burrows in dead wood; hollow stems (e.g. <em>Phragmites<\/em>, bamboo) and straw tubes. <strong>Nesting material:<\/strong> Cell partitions and nest plug are made of mud. (Kitamura and Maeta, 1969; Maeta, 1969, 1978)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Polylectic; pollen sources include plants belonging to 9 different families; flowers containing no or very little nectar, such as <em>Juglans<\/em> (Juglandaceae), <em>Quercus<\/em> (Fabaceae) or <em>Liquidambar<\/em> (Altingiaceae), are important pollen hosts (Quest, 2009; Haider et al., 2013).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Osmia) tricornis <\/em><\/strong><span style=\"font-weight: normal\">Latreille, 1811<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1811 <em>Osmia <strong>tricornis<\/strong><\/em> Latreille, Encyclop\u00e9die M\u00e9thodique, Histoire Naturelle, Insectes, vol. 8, p. 575. <span style=\"text-decoration: underline\">Type material<\/span>: Syntypes m(m), f(f), &#8220;environs d&#8217;Aix, Montpellier&#8221; [France].-Combination <em>Osmia (Ceratosmia) tricornis<\/em> Latreille<!-- [if supportFields]&gt;&lt;![endif]--> in Schmiedeknecht (1885: 19 [885]). Combination <em>Osmia (Pachyosmia) tricornis<\/em> Latreille in Friese (1911b: 126).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe:<\/strong> AND, E, E(Ba), <span style=\"text-decoration: underline\">F<\/span>, F(Co), I, I(Sa), P. <strong>Northern Africa:<\/strong> DZ, E(Ca), LAR, MA, TN.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Benoist (1931); Ducke (1900); Peters (1978a); Radoszkowski (1887a: 292); Schmiedeknecht (1885-1886).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: insect burrows in dead wood and drilled borings in wooden blocks; hollow stems (e.g. <em>Phragmites<\/em>); insect burrows in the ground; empty snail shell (<em>Cepaea<\/em>) in the fissure of a stone wall; abandoned cells in exposed nests of <em>Chalicodoma<\/em>. <strong>Nesting material: <\/strong>Cell partitions and nest plug are made of mud. (Benoist, 1931; O&#8217;Toole and Raw, 1991; Vicens et al., 1993, 1994; A. M\u00fcller, personal observation)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Flower preferences: <\/strong>Polylectic; pollen sources include plants belonging to 12 different families; flowers of Cistaceae and to a lesser extent <em>Papaver<\/em> (Papaveraceae) are the two most important pollen hosts, both of which contain no or very little nectar (Vicens et al., 1994; Haider et al., 2013).<\/p>\n<h3><strong><span style=\"text-decoration: underline\"><em>emarginata<\/em> species group<\/span> (&#8220;<em>Aceratosmia<\/em>&#8220;)<\/strong><\/h3>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Osmia) emarginata <\/em><\/strong><span style=\"font-weight: normal\">Lepeletier, 1841<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1841 <em>Osmia <strong>emarginata<\/strong><\/em> Lepeletier, Histoire Naturelle des Insectes, Hym\u00e9nopt\u00e8res, vol. 2, p. 317 [not seen]. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Bois du V\u00e9sinet, pr\u00e8s Saint-Germain-en-Laye&#8221; [France], MNHN (Paris). <span style=\"text-decoration: underline\">Type species<\/span> of <em>Aceratosmia<\/em> Schmiedeknecht.-Combination <em>Osmia (Aceratosmia) emarginata<\/em> Lepeletier in Schmiedeknecht (1885: 19 [885]). Combination <em>Osmia (Pachyosmia) emarginata<\/em> Lepeletier in Friese (1911b: 125).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>E, <span style=\"text-decoration: underline\">F<\/span>, F(Co), I, I(Sa), P, <del>SCG<\/del><em>. <\/em><strong>Northern Africa: <\/strong>DZ, MA<em>, <\/em>TN<em>.<\/em><strong> Southwestern Asia: <\/strong><span style=\"text-decoration: line-through\">IL<\/span><em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Banaszak and Romasenko (2001); Benoist (1931); Ducke (1900, incl. <em>Osmia mustelina<\/em>); Peters (1978a); Schmiedeknecht (1885-1886, incl. <em>Osmia mustelina<\/em>); Tkalcu (1971). <strong>Misapplication:<\/strong> Historically often confounded with <em>Osmia mustelina<\/em> Gerstaecker, see Tkalcu (1971).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: fissures between stones and cavities in rocks and stones; metal tubes. <strong>Nesting material: <\/strong>The cells, which are built side by side, are entirely built from chewed leaves. The outermost layer of the nest consists of empty cells. Often, several females communally nest in the same fissure or cavity. <strong>Cleptoparasite:<\/strong> <em>Stelis franconica<\/em>. (Banaszak and Romasenko, 2001; Benoist, 1931; Deleurance, 1949; Ferton, 1901a; Genoud and Dufr\u00eane, 2017; Tkalcu, 1971; Weigand, 2010, as <em>Osmia mustelina<\/em>)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Flower preferences: <\/strong>Polylectic; known pollen sources are <em>Echium<\/em> (Boraginaceae), Cistaceae, Fabaceae, <em>Papaver<\/em> (Papaveraceae) and Lamiaceae (Haider et al., 2013).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>emarginata <\/strong><\/em><span style=\"font-weight: normal\">Lepeletier, 1841<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1841 Automatically established nominotypical subspecific taxon (original description and type specimen are identical with those of the species rank taxon of the same name above).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>E, <span style=\"text-decoration: underline\">F<\/span>, F(Co), I, I(Sa), P, <del>SCG<\/del><em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Peters (1978a).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>infuscata <\/strong><\/em><span style=\"font-weight: normal\">Peters, 1978<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1928 <em>Osmia emarginata <strong>fusca<\/strong> <\/em>Benoist, Bulletin de la Soci\u00e9t\u00e9 des Sciences Naturelles du Maroc (Rabat), 8: 215. NOMEN PRAEOCCUPATUM [not <em>Apis fusca<\/em> Christ, 1791; not <em>Osmia fusca<\/em> Gistel, 1857]. <span style=\"text-decoration: underline\">Type material<\/span>: Syntypes m(m), f(f), \u201eMoyen Atlas, au sud de Taza: Tizi n\u2019Bouft\u00e8ne (2300 m), Tasrah des Ighezrane (2300 m)\u201c [Morocco], MNHN (Paris).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1978 <em>Osmia emarginata <strong>infuscata<\/strong><\/em> Peters, Senckenbergiana Biologica (Frankfurt), 58: 310. NOMEN NOVUM with same type specimen for preoccupied <em>Osmia emarginata fusca<\/em> Benoist, 1928 [not <em>Apis fusca<\/em> Christ, 1791; not <em>Osmia fusca<\/em> Gistel, 1857].<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Africa: <\/strong>DZ, <span style=\"text-decoration: underline\">MA<\/span>, TN.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Tkalcu (1971).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Osmia) fedtschenkoi <\/em><\/strong><span style=\"font-weight: normal\">Morawitz, 1875<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1875 <em>Osmia <strong>Fedtschenkoi<\/strong><\/em> Morawitz, Travel to Turkestan by A. P. Fedtschenko, p. 81. <span style=\"text-decoration: underline\">Type material<\/span>: Syntypes m(m), f(f), &#8220;prope Samarkand&#8221; [Uzbekistan].-Combination <em>Osmia (Chalcosmia) fedtschenkoi <\/em>Morawitz in Friese (1911b: 114). Combination <em>Osmia (Aceratosmia) fedtschenkoi<\/em> Morawitz in Tkalcu (1969a: 340).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1904 <em>Osmia<\/em> <em>(Ceratosmia)<\/em> <strong><em>balucha<\/em><\/strong> Nurse, Journal of the Bombay Natural History Society, 15: 573. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype f, by designation of Tkalcu (1969a: 341), &#8220;Quetta&#8221; [Pakistan], BMNH (London); paralectotype m.-Synonymy in Tkalcu (1969a: 340).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1930 <em>Osmia<\/em> <strong><em>usbekistana<\/em><\/strong> Cockerell, The Annals and Magazine of Natural History, ser. 10, 5: 408. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype f, by designation of Tkalcu (1969a: 341), &#8220;Tashkent&#8221; [Uzbekistan], BMNH (London).-Synonymy in Tkalcu (1969a: 340).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Asia: <\/strong>KZ, TJ, TM, <span style=\"text-decoration: underline\">UZ<\/span><em>. <\/em><strong>Southwestern Asia: <\/strong>AFG, PAK<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Ducke (1900); Peters (1978a); Tkalcu (1969a: 341-342).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: insect burrows in dead wood and drilled borings in wooden blocks; hollow stems (e.g. <em>Arundo<\/em>); insect burrows in the soil (e.g. of <em>Anthophora<\/em>). <strong>Nesting material: <\/strong>Cell partitions and nest plug are made of mud. In larger cavities, cells are irregularly arranged with cell walls partially or entirely built of mud. (Beiko, 1987; Nurse, 1904; Ponomareva, 1958)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Polylectic with a strong preference for Fabaceae; other known pollen sources are Rosaceae, Brassicaceae, <em>Lonicera<\/em> (Caprifoliaceae) and <em>Salix<\/em> (Salicaceae) (Haider et al., 2013).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Osmia) melanocephala <\/em><\/strong><span style=\"font-weight: normal\">Morawitz, 1875<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1875 <em>Osmia <strong>melanocephala<\/strong><\/em> Morawitz, Travel to Turkestan by A. P. Fedtschenko, p. 80. <span style=\"text-decoration: underline\">Type material<\/span>: Syntypes m(m), f(f), &#8220;in valle Sarafschan&#8221; [Tajikistan].-Combination <em>Osmia (Pachyosmia) melanocephala<\/em> Morawitz in Friese (1911b: 124). Combination <em>Osmia (Aceratosmia) melanocephala<\/em> Morawitz in Tkalcu (1969a: 340).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Asia: <\/strong>KS, KZ, MGL, RC, <span style=\"text-decoration: underline\">TJ<\/span>, TM, UZ; <del>RUS<\/del><em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Ducke (1900); Peters (1978a); Radoszkowski (1886: 13); Tkalcu (1995: 142); Wu (2006); Zanden (1996b: 895).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Flower preferences: <\/strong>Polylectic with a strong preference for Fabaceae; other known pollen sources are Rosaceae, Monokots, <em>Lonicera<\/em> (Caprifoliaceae) and Boraginaceae (Haider et al., 2013).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Osmia) mustelina <\/em><\/strong><span style=\"font-weight: normal\">Gerstaecker, 1869<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1869 <em>Osmia <strong>mustelina<\/strong><\/em> Gerstaecker, Entomologische Zeitung (Stettin), 30: 348. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype f, by designation of Tkalcu (1971: 225), &#8220;Franken (bei Culmbach)&#8221; [Germany], ZMHB (Berlin); paralectotype f.-Combination <em>Osmia (Aceratosmia) mustelina<\/em> Gerstaecker in Tkalcu (1969a: 340).-Synonymy with <em>Osmia emarginata<\/em> Lepeletier in Friese (1911b: 125), rejected by Tkalcu (1971).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>A, ARM, AZ, B, BG, BIH, CH, CZ, <span style=\"text-decoration: underline\">D<\/span>, FL, GE, GR, GR(Aegean Islands), H, HR, I, I(Si), L, MK, PL, RO, RUS(SR), SCG, SK, SLO, UA(Crimea). <strong>Southwestern Asia: <\/strong>IL, IR, RL, TR<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Amiet et al.(2004); Banaszak and Romasenko (2001); Ducke (1900, incl. <em>Osmia emarginata<\/em>); Medvedeva (1978, as <em>Osmia emarginata<\/em>); M\u00f3cz\u00e1r (1958, as <em>Osmia emarginata<\/em>); Peters (1978a); Scheuchl (1996); Schmiedeknecht (1885-1886, incl. <em>Osmia emarginata<\/em>); Tkalcu (1971; 1974b: 339-340). <strong>Misapplication:<\/strong> Historically often confounded with <em>Osmia emarginata<\/em> Lepeletier, see Tkalcu (1971).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: fissures between stones and cavities in rocks, stones and walls; rarely in hollow stems or in abandoned cells of <em>Megachile (Chalicodoma) parietina.<\/em> <strong>Nesting material:<\/strong> The cells, which are built side by side, entirely consist of chewed leaves (e.g. of <em>Helianthemum, Cytisus<\/em>). While in larger cavities the entire cells are built from leaf pulp, only the side walls are built in narrow cavities. The outermost layer of the nest consists of empty cells. Often, several females communally nest in the same fissure or cavity. (Banaszak and Romasenko, 2001; Bonelli, 1967; Friese, 1895, 1923; Grandi, 1964; M\u00fcller et al., 1997; Stoeckhert, 1933; Westrich, 1989)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Flower preferences: <\/strong>Polylectic; known pollen sources are Fabaceae, <em>Papaver<\/em> (Papaveraceae), Cistaceae, Boraginaceae (<em>Echium, Anchusa<\/em>) and four other plant families (Amiet et al., 2004; Westrich, 1989; Haider et al., 2013).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>griseohirta <\/strong><\/em><span style=\"font-weight: normal\">Alfken, 1935<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1935 <em>Osmia emarginata<strong> griseohirta<\/strong><\/em> Alfken, Mitteilungen aus dem Entomologischen Verein in Bremen, 23: 23. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype m, by designation of Tkalcu (1971), &#8220;Elburs, Pish-Kuh&#8221; [Iran], ZMHB (Berlin).-Combination <em>Osmia (Osmia) mustelina griseohirta<\/em> Alfken in Tkalcu (1971: 226).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe:<\/strong> ARM, AZ, GE, GR. <strong>Southwestern Asia:<\/strong> <span style=\"text-decoration: underline\">IR<\/span>, TR.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Peters (1978a); Tkalcu (1971).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>mustelina <\/strong><\/em><span style=\"font-weight: normal\">Gerstaecker, 1869<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1869 Automatically established nominotypical subspecific taxon (original description and type specimen are identical with those of the species rank taxon of the same name above).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>A, B, CH, BG, BIH, CZ, <span style=\"text-decoration: underline\">D<\/span>, FL, GR, H, HR, I, L, MK, PL, RUS, SCG, SK, SLO, UA(Crimea)<em>. <\/em><strong>Southwestern Asia: <\/strong>IL, RL, TR<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Peters (1978a).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>umbrosa <\/strong><\/em><span style=\"font-weight: normal\">Peters, 1978<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1978 <em>Osmia (Osmia) mustelina <strong>umbrosa<\/strong><\/em> Peters, Senckenbergiana Biologica (Frankfurt), 58: 313. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Palermo&#8221; [Italy: Sicily], SMFD (Frankfurt); paratypes ff.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>I, <span style=\"text-decoration: underline\">I(Si)<\/span><em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known.<\/p>\n<h3><strong><em>Osmia (Osmia) mutensis<\/em><\/strong><strong> <\/strong><span style=\"font-weight: normal\">Peters, 1978<\/span><\/h3>\n<p style=\"text-align: left;padding-left: 30px\">1978 <em>Osmia (Osmia) <strong>mutensis<\/strong><\/em> Peters, Senckenbergiana Biologica (Frankfurt), 58: 315. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Mut, Sertavul, 1600 m&#8221; [Turkey], SMFD (Frankfurt).<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Distribution-Southwestern Asia:<\/strong><em> <\/em><span style=\"text-decoration: underline\">TR<\/span><em>.<\/em><\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Identification-<\/strong><strong>Keys, Descriptions:<\/strong><strong> <\/strong>No supplementary or more detailed morphological description known. <strong>Note: <\/strong>Male unknown.<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"text-align: left;padding-left: 30px\"><strong>Flower preferences: <\/strong>Unknown.<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Osmia) nigrohirta <\/em><\/strong><span style=\"font-weight: normal\">Friese, 1899<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1899 <em>Osmia bicolor <strong>nigrohirta<\/strong><\/em> Friese, Entomologische Nachrichten (Berlin), 25: 25. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Poros&#8221; [Greece], ZMHB (Berlin).-Combination <em>Osmia (Aceratosmia) nigrohirta<\/em> Friese in Tkalcu (1969a: 340).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>AL, ARM, AZ, <span style=\"text-decoration: underline\">GR<\/span>, MK; Caucasus. <strong>Northern Asia:<\/strong> TM. <strong>Southwestern Asia: <\/strong>IL, IR, RL, TR<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Ducke (1900); Peters (1978a); Tkalcu (1974b: 339-340).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Polylectic; known pollen sources are Fabaceae (e.g.<em> Onobrychis<\/em>), <em>Echium<\/em> (Boraginaceae), Lamiaceae, Cistaceae, Asteraceae, Brassicaceae and four other plant families (Haider et al., 2013).<\/p>\n","protected":false},"excerpt":{"rendered":"<p>The subgenus Osmia Panzer is distributed both in the Palaearctic and the Nearctic region. It contains 27 described species, 25 of which occur in the Palaearctic. The North American species Osmia lignaria Say, which has been introduced at one locality in Spain (Ortiz-S\u00e1nchez, 2011), is not listed below. Due to the close morphological resemblance, the [&hellip;]<\/p>\n","protected":false},"author":1092,"featured_media":0,"parent":46,"menu_order":0,"comment_status":"open","ping_status":"closed","template":"","meta":{"footnotes":""},"class_list":["post-2683","page","type-page","status-publish","hentry"],"_links":{"self":[{"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/pages\/2683","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/pages"}],"about":[{"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/types\/page"}],"author":[{"embeddable":true,"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/users\/1092"}],"replies":[{"embeddable":true,"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/comments?post=2683"}],"version-history":[{"count":7,"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/pages\/2683\/revisions"}],"predecessor-version":[{"id":7425,"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/pages\/2683\/revisions\/7425"}],"up":[{"embeddable":true,"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/pages\/46"}],"wp:attachment":[{"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/media?parent=2683"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}