{"id":162,"date":"2009-01-22T00:08:52","date_gmt":"2009-01-21T23:08:52","guid":{"rendered":"https:\/\/wpethzprd.ethz.ch\/osmiini\/?page_id=162"},"modified":"2025-12-28T16:25:42","modified_gmt":"2025-12-28T15:25:42","slug":"pyrosmia","status":"publish","type":"page","link":"https:\/\/blogs.ethz.ch\/osmiini\/palaearctic-species\/osmia\/pyrosmia\/","title":{"rendered":"Subgenus Pyrosmia"},"content":{"rendered":"<p style=\"text-align: left\">The subgenus <em>Pyrosmia<\/em> Tkalcu is confined to the Palaearctic region. It contains 33 described species.<\/p>\n<p style=\"text-align: left\"><a href=\"https:\/\/blogs.ethz.ch\/osmiini\/files\/2009\/09\/osmia-ferruginea-nico_web.jpg\"><img loading=\"lazy\" decoding=\"async\" class=\"alignnone size-full wp-image-2856\" src=\"https:\/\/blogs.ethz.ch\/osmiini\/files\/2009\/09\/osmia-ferruginea-nico_web.jpg\" alt=\"\" width=\"433\" height=\"296\" \/><\/a><\/p>\n<h5 style=\"text-align: left\"><span style=\"font-weight: normal\">Female of <\/span><em><span style=\"font-weight: normal\">Osmia (Pyrosmia) ferruginea<\/span><\/em><span style=\"font-weight: normal\"> Latreille, 1811. Foto N. Vereecken.<\/span><\/h5>\n<h3 style=\"text-align: left\">Species accounts<\/h3>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) amathusica <\/em><\/strong><span style=\"font-weight: normal\">Mavromoustakis, 1937<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1937 <em>Osmia (Chalcosmia) <strong>amathusica<\/strong><\/em> Mavromoustakis, Annals and Magazine of Natural History (London), 20: 522. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Amathus (ruins)&#8221; [Cyprus], DAAN (Nicosia); paratypes mm, ff.-Combination <em>Osmia (Diceratosmia) amathusica<\/em> Mavromoustakis in Tkalcu (1969a: 333). Combination <em>Osmia (Caerulosmia) amathusica<\/em> Mavromoustakis in Zanden (1988b: 123).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution- Europe: <\/strong>AZ, GR, GR(Cr), I(Si)<em>. <\/em><strong>Southwestern Asia:<\/strong> <span style=\"text-decoration: underline\">CY<\/span>, IL, JOR, RL, SYR, TR.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Mavromoustakis (1956a: 857); Warncke (1992b); Zanden (1991a).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: crevice in a stone containing two cells. <strong>Nesting material: <\/strong>The cells are constructed with very small pebbles and chewed leaves. (Mavromoustakis, 1948c)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences:<\/strong> The only three pollen samples available so far (from three different localities) consist of pollen of Fabaceae (e.g. <em>Trifolium<\/em>) and Lamiaceae (A. M\u00fcller, unpublished).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) cyanoxantha <\/em><\/strong><span style=\"font-weight: normal\">P\u00e9rez, 1879<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1879 <em>Osmia <strong>cyanoxantha<\/strong><\/em> P\u00e9rez, Actes de la Soci\u00e9t\u00e9 Linn\u00e9enne de Bordeaux, 33: 185. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype f, by designation of Zanden (1991a: 52), &#8220;Marseille&#8221; [France], MNHN (Paris); paralectotypes mm, ff.-Combination <em>Osmia (Chalcosmia) cyanoxantha<\/em> P\u00e9rez in Schmiedeknecht (1885: 20 [886]). Combination <em>Osmia (Diceratosmia) cyanoxantha<\/em> P\u00e9rez in Tkalcu (1969a: 333). Combination <em>Osmia (Caerulosmia) cyanoxantha<\/em> P\u00e9rez in Zanden (1988b: 123).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1991 <em>Osmia (Caerulosmia)<\/em> <strong><em>cyanoxura<\/em><\/strong> Zanden, Linzer Biologische Beitr\u00e4ge, 23: 53. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype m, &#8220;Erimanthos Mts., Pelop.&#8221; [Greece], RMNH (Leiden); paratypes mm, ff.-Synonymy in Warncke (1992b: 909).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>ARM, AZ, BG, E?, <span style=\"text-decoration: underline\">F<\/span>, GR, GR(Aegean Islands), HR, I, I(Si), MK, P?, RUS(SR), SLO, UA(Crimea). <strong>Northern Africa: <\/strong>DZ, MA, TN.<strong> Southwestern Asia: <\/strong>CY, IL, IR, JOR, RL, SYR, TR<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Amiet et al. (2004); Benoist (1931); Ducke (1900); Schmiedeknecht (1885-1886); Warncke (1992b); Zanden (1991a).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>This species appears to use both preexisting cavities and stone surfaces as nesting sites. Preexisting cavities: abandoned cells in exposed nests of <em>Megachile (Chalicodoma) parietina<\/em>; small cavities and holes in rocks or stones containing mostly 1, rarely up to 4 cells. Exposed nests: upper surface of a stone with 2-3 cells. <strong>Nesting material: <\/strong>Cell partitions are made of chewed leaves intermixed with small pebbles or occasionally of chewed leaves only. The nest plug is made of two to four layers of small pebbles (1-2mm in diameter) cemented together with leaf pulp, its convex surface expanding 2-3mm to the adjacent rock surface. Cells of exposed nests are entirely constructed from very small pebbles cemented together with chewes leaves and eventually covered by an additional layer of the same material. (Benoist, 1931; Ferton, 1901a, 1914, 1920; Mavromoustakis, 1948c)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Flower preferences: <\/strong>Polylectic; pollen sources recorded so far include Fabaceae (e.g. <em>Lotus<\/em>), <em>Echium<\/em> (Boraginaceae) and Lamiaceae (A. M\u00fcller, unpublished, based on 10 pollen samples from 8 different localities).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) derasa <\/em><\/strong><span style=\"font-weight: normal\">P\u00e9rez, 1895<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1895 <em>Osmia <strong>derasa<\/strong><\/em> P\u00e9rez, Esp\u00e8ces nouvelles de mellif\u00e8res de Barbarie, p. 12. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype f, by designation of Tkalcu (1970: 9), &#8220;T\u00e8niet&#8221; [Algeria], MNHN (Paris).-Combination <em>Osmia (Diceratosmia) derasa<\/em> P\u00e9rez in Tkalcu (1969a: 333). Combination <em>Osmia (Caerulosmia) derasa<\/em> P\u00e9rez in Zanden (1988b: 123).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1929 <em>Osmia <strong>Crosi<\/strong><\/em> Benoist, Bulletin de la Soci\u00e9t\u00e9 Entomologique de France, 1929: 96. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype m, by designation of Zanden (1985: 55), &#8220;Mascara&#8221; [Algeria], MNHN (Paris); paralectotype m.-Combination <em>Osmia (Diceratosmia) crosi<\/em> Benoist in Tkalcu (1969a: 333). Combination <em>Osmia (Caerulosmia) crosi<\/em> Benoist in Zanden (1988b: 123).-Synonymy with <em>Osmia derasa<\/em> P\u00e9rez in Warncke (1992b: 905). Synonymy with <em>Osmia tricolor<\/em> Saunders in Zanden (1995: 429), which is a mistake (T. Griswold, personal communication).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1950 <em>Osmia<\/em> <strong><em>pseudoderasa<\/em><\/strong> Benoist, Bulletin de la Soci\u00e9t\u00e9 des Sciences Naturelles du Maroc (Rabat), 30: 184. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Midelt&#8221; [Morocco], MZLS (Lausanne).-Synonymy in Zanden (1985: 68).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Africa:<\/strong> <span style=\"text-decoration: underline\">DZ<\/span>, LAR, MA, TN.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Tkalcu (1970: 9; 1975a: 183); Zanden (1991a; 1994b: 1120).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong><strong>Nesting biology-Nesting site: <\/strong><\/strong>Preexisting cavities: abandoned cells in exposed nests of <em>Hoplitis (Hoplitis). <\/em><strong><strong>Nesting material:<\/strong> <\/strong>Unknown. (C. Sedivy, C. Praz and A. M\u00fcller, personal observation)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Flower preferences:<\/strong> Polylectic; pollen sources recorded so far include Fabaceae and <em>Echium<\/em> (Boraginaceae) (A. M\u00fcller, unpublished, based on 5 pollen samples from 5 different localities).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>derasa <\/strong><\/em><span style=\"font-weight: normal\">P\u00e9rez, 1896<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1896 Automatically established nominotypical subspecific taxon (original description and type specimen are identical with those of the species rank taxon of the same name above).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Africa: <\/strong><span style=\"text-decoration: underline\">DZ<\/span>, LAR, MA, TN<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions:<\/strong>No supplementary or more detailed morphological description known.<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>violetta <\/strong><\/em><span style=\"font-weight: normal\">Warncke, 1992<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1992 <em>Osmia (Pyrosmia) derasa <strong>violetta<\/strong><\/em> Warncke, Linzer Biologische Beitr\u00e4ge, 24: 905. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Cap Bon&#8221; [Tunisia], OLML (Linz); paratypes m(m), f(f).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Africa: <\/strong><span style=\"text-decoration: underline\">TN<\/span><em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known.<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) dilaticornis <\/em><\/strong><span style=\"font-weight: normal\">Morawitz, 1875<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1875 <em>Osmia <strong>dilaticornis<\/strong><\/em> Morawitz, Travel to Turkestan by A. P. Fedtschenko, p. 87. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype m, &#8220;prope Taschkent&#8221; [Uzbekistan], ZMUM (Moscow).-Combination <em>Osmia (Diceratosmia) dilaticornis<\/em> Morawitz in Tkalcu (1969a: 333). Combination <em>Osmia (Caerulosmia) dilaticornis<\/em> Morawitz in Zanden (1988b: 123).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>AZ, GR, GR(Aegean Islands)<em>. <\/em><strong>Northern Asia:<\/strong> KZ, TJ, <span style=\"text-decoration: underline\">UZ<\/span>. <strong>Southwestern Asia:<\/strong> IL, IR, JOR, RL, SYR, TR.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Warncke (1992b); Zanden (1989: 85-86, 1991a).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<strong><br \/>\n<\/strong><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Possibly oligolectic on Fabaceae (A. M\u00fcller, unpublished, based on 7 pollen samples from 4 different localities).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) <\/em><i>elbaba<\/i><\/strong><i style=\"font-weight: bold\"> <\/i><span style=\"font-weight: normal\">Warncke, 1992<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1992 <em>Osmia (Pyrosmia)<\/em> <strong><em>elbaba<\/em><\/strong> Warncke, Linzer Biologische Beitr\u00e4ge, 24: 904. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Medjez el Bab&#8221; [Tunisia], OLML (Linz); paratype f.-Synonymy with <em>Osmia cyanoxantha<\/em> in Zanden (1996b: 884), rejected by A. M\u00fcller (unpublished).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-<\/strong><strong>Northern Africa: <\/strong>DZ, MA, TN.<strong> <\/strong><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: empty snail shells with 3 brood cells. <strong>Nesting material: <\/strong>Cell partitions and nest plug are made of chewed leaves. (Ferton, 1914, as <em>Osmia leucopyga<\/em>)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Flower preferences: <\/strong>Unknown.<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) ferruginea <\/em><\/strong><span style=\"font-weight: normal\">Latreille, 1811 <\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1811 <em>Osmia <strong>ferruginea<\/strong><\/em> Latreille, Encyclop\u00e9die M\u00e9thodique, Histoire Naturelle, Insectes, vol. 8, p. 579. <span style=\"text-decoration: underline\">Type material<\/span>: f(f), [Egypt]. <span style=\"text-decoration: underline\">Type species<\/span> of <em>Pyrosmia<\/em> Tkalcu.-Combination <em>Osmia (Chalcosmia) ferruginea <\/em>Latreille in Friese (1911b: 109).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1849 <em>Osmia<\/em> <strong><em>metallica<\/em><\/strong> Lucas, Exploration scientifique de l&#8217;Alg\u00e9rie, p. 191. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;environs de Misserghin&#8221; [Algeria], MNHN (Paris).-Synonymy in Zanden (1988b: 127).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>E, E(Ba), F, F(Co), I(Sa), I(Si), M, P; GR?<em>. <\/em><strong>Northern Africa: <\/strong>DZ, <span style=\"text-decoration: underline\">ET<\/span>, LAR, MA, TN<em>. <\/em><strong>Southwestern Asia:<\/strong> CY, IL, JOR, SYR.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Benoist (1931); Ducke (1900); Warncke (1992b).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: empty snail shells of small to large size (e.g. <em>Bulimus, Cerithium, Cernuella, Cochlicella, Eobania, Helicella, Helix, Iberellus, Otala, Rumina, Sphincterochila, Theba, Trochochlea<\/em>) with 1-10 cells per shell; hollow stems (e.g. <em>Phragmites<\/em>). <strong>Nesting material:<\/strong> Cell partitions and nest plug are made of chewed leaves (e.g. of <em>Convolvulus, Lavathera<\/em>). Occasionally, small pebbles or other particles are incorporated into the leaf matrix of the nest plug. The space between the partition of the outermost cell and the nest plug is sometimes loosely filled with small pebbles, earth crumbs, seeds or leaflets. The closed shells are neither moved nor covered with patches of leaf pulp. <strong>Brood parasites:<\/strong> <em>Sapyga quinquepunctata<\/em> (Sapygidae), <em>Anthrax aethiops<\/em> (Bombyliidae). (Alfken, 1914; Bogusch et al., 2020; Benoist, 1931; Ferton, 1890, 1897, 1905; Grandi, 1961; Haeseler, 1997; Mavromoustakis, 1939b, 1952; Moreno-Rueda et al., 2008; Saunders, 1908; G. Le Goff, personal observation; C. Sedivy, C. Praz and A. M\u00fcller, personal observation)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Polylectic with a preference for Fabaceae; additional pollen sources include Boraginaceae (<em>Echiochilon, Echium<\/em>), <em>Reseda<\/em> (Resedaceae), Lamiaceae, Antirrhineae, Brassicaceae, Cistaceae and monocots (Bogusch et al., 2020; A. M\u00fcller, unpublished, based on 64 pollen samples from 41 different localities and on field observations).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>ferruginea <\/strong><\/em><span style=\"font-weight: normal\">Latreille, 1811<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1811 Automatically established nominotypical subspecific taxon (original description and type specimen are identical with those of the species rank taxon of the same name above).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>E, F, P; GR<em>.<\/em><strong>Northern Africa: <\/strong>DZ, <span style=\"text-decoration: underline\">ET<\/span>, LAR, MA, TN<em>. <\/em><strong>Southwestern Asia:<\/strong> CY, IL, JOR.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known.<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>igneopurpurea <\/strong><\/em><span style=\"font-weight: normal\">Costa, 1882<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1882 <em>Osmia <strong>igneo-purpurea<\/strong><\/em> Costa, Rendiconti dell&#8217;Academia delle Scienze Fisiche e Matematiche de Napoli, 21: 198. <span style=\"text-decoration: underline\">Type material<\/span>: f(f), [Italy: Sardinia].-Synonymy with <em>Osmia ferruginea<\/em> Latreille in Tkalcu (1975a: 182).-Combination <em>Osmia (Pyrosmia) ferruginea igneopurpurea<\/em> Costa in Warncke (1992b: 917).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>F(Co), <span style=\"text-decoration: underline\">I(Sa)<\/span>, I(Si), M<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions:<\/strong>Benoist (1931);Ferton (1897: 39); Schmiedeknecht (1885-1886); Warncke (1992b).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) forticornis <\/em><\/strong><span style=\"font-weight: normal\">Zanden, 1989 <\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1989 <em>Osmia (Caerulosmia) <strong>forticornis<\/strong><\/em> Zanden, Entomologische Abhandlungen (Dresden), 53: 85. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype m, &#8220;Chelmos Pelopon., 1900 m&#8221; [Greece], RMNH (Leiden); paratypes mm, ff.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>ARM, AZ, <span style=\"text-decoration: underline\">GR,<\/span><em> <\/em>GR(Aegean Islands). <strong>Southwestern Asia: <\/strong>RL, TR<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Warncke (1992b); Zanden (1991a).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Polylectic; pollen sources recorded so far include Fabaceae and Lamiaceae (A. M\u00fcller, unpublished, based on 5 pollen samples from 4 different localities).<\/p>\n<h3 style=\"text-align: left\"><strong><em><span style=\"font-style: normal;font-weight: normal\"><strong><em>Osmia (Pyrosmia) gemmea <\/em><\/strong>P\u00e9rez, 1896<\/span><\/em><\/strong><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1895 <em>Osmia<\/em> <strong><em>purpurea<\/em><\/strong> P\u00e9rez, Esp\u00e8ces nouvelles de mellif\u00e8res de Barbarie, p. 11. NOMEN PRAEOCCUPATUM [not <em>Osmia purpurea <\/em>Cresson, 1864]. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype m, by designation of Zanden (1991a: 57), &#8220;Tunis&#8221; [Tunisia], MNHN (Paris); paralectotype f.<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1896 <em>Osmia <strong>gemmea<\/strong><\/em> P\u00e9rez, Correctures: Esp\u00e8ces nouvelles de mellif\u00e8res de Barbarie, not paginated single page appendix to P\u00e9rez (1895). NOMEN NOVUM with same type specimen for preoccupied <em>Osmia purpurea<\/em> P\u00e9rez, 1895 [not <em>Osmia purpurea <\/em>Cresson, 1864].-Combination <em>Osmia (Chalcosmia) gemmea<\/em> P\u00e9rez in Friese (1911b: 109). Combination <em>Osmia (Diceratosmia) gemmea<\/em> P\u00e9rez in Tkalcu (1969a: 333). Combination <em>Osmia (Caerulosmia) gemmea<\/em> P\u00e9rez in Zanden (1988b: 123).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1899 <em>Osmia<\/em> <strong><em>ligulicornis<\/em><\/strong> Friese, Entomologische Nachrichten (Berlin), 25: 330. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype f, by designation of Zanden (1987: 78), &#8220;Jaffa&#8221; [Israel], ZMHB (Berlin).-Synonymy in Zanden (1987: 78).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Africa: <\/strong>DZ, ET, LAR, MA, <span style=\"text-decoration: underline\">TN<\/span><em>. <\/em><strong>Southwestern Asia: <\/strong>IL<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Alfken (1942: 214-215); Warncke (1992b); Zanden (1991a).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: empty snail shells (e.g. <em>Trochoidea<\/em>). <strong>Nesting material: <\/strong>Unknown. (Alfken, 1942; Morice, 1900; Zanden, 1991a)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Polylectic; pollen sources recorded so far are Fabaceae (e.g. <em>Retama<\/em>), <em>Echium <\/em>(Boraginaceae),<em> Reseda<\/em> (Resedaceae) and probably Antirrhineae (A. M\u00fcller, unpublished, based on 8 pollen samples from 7 different localities).<\/p>\n<h3><strong><em>Osmia (Pyrosmia) hermona <\/em><\/strong><span style=\"font-weight: normal\">Warncke, 1992<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1992 <em>Osmia (Pyrosmia) <strong>hermona<\/strong><\/em> Warncke, Linzer Biologische Beitr\u00e4ge, 24: 906. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype m, &#8220;Mt. Hermon in 2050 m&#8221; [Israel], OLML (Linz); paratypes mm, ff.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Asia: <\/strong>TJ<em>. <\/em><strong>Southwestern Asia: <\/strong><span style=\"text-decoration: underline\">IL<\/span>, SYR, TR<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Unknown. The only pollen sample available so far consists of pollen of Fabaceae (A. M\u00fcller, unpublished).<strong><br \/>\n<\/strong><\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) laticauda <\/em><\/strong><span style=\"font-weight: normal\">Stanek, 1969<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1969 <em>Osmia (Chalcosmia) <strong>laticauda<\/strong><\/em> Stanek, Nachrichten des Naturwissenschaftlichen Museums der Stadt Aschaffenburg, 78: 5. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype m, &#8220;Mut-Sertavul, 1410 m&#8221; [Turkey], SMFD (Frankfurt).-Combination <em>Osmia (Caerulosmia) laticauda<\/em> Stanek in Zanden (1988b: 123).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1992 <em>Osmia <strong>tichodroma<\/strong><\/em> Warncke, Linzer Biologische Beitr\u00e4ge, 24: 912. NOMEN NOVUM with same type specimen for preoccupied <em>Osmia laticauda<\/em> Stanek, 1969 [not <em>Heriades laticaudus<\/em> Benoist, 1938].<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe:<\/strong> BG, GR, GR(Aegean Islands), MK<em>. <\/em><strong>Southwestern Asia: <\/strong>IR, IRQ, <span style=\"text-decoration: underline\">TR<\/span><em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Warncke (1992b); Zanden (1991a).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Polylectic; pollen sources recorded so far include Fabaceae and Lamiaceae (A. M\u00fcller, unpublished, based on 5 pollen samples from 4 different localities).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) leucopyga <\/em><\/strong><span style=\"font-weight: normal\">Ducke, 1899<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1899 <em>Osmia<\/em> <strong><em>leucopyga<\/em><\/strong> Ducke, Entomologische Nachrichten (Berlin), 25: 214. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Constantine&#8221; [Algeria], ZMHB (Berlin).-Combination <em>Osmia (Caerulosmia) leucopyga <\/em>Ducke in Zanden (1988b: 123).-Synonymy with <em>Osmia lobata<\/em> Friese, 1899 in Warncke (1992b: 910), rejected by A. M\u00fcller (unpublished).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe:<\/strong> E, P. <strong>Northern Africa:<\/strong> <span style=\"text-decoration: underline\">DZ<\/span>, MA, TN?.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Ducke (1900); Zanden (1991a).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: <span lang=\"EN-US\">abandoned brood cells of <i>Megachile<\/i> (<i>Chalicodoma<\/i>), burrows in the soil. <strong>Nesting material:<\/strong> Unknown. (Zanden, 1991b).<\/span><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting material: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Unknown.<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) lobata <\/em><\/strong><span style=\"font-weight: normal\">Friese, 1899<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1899 <em>Osmia <strong>lobata<\/strong><\/em> Friese, Entomologische Nachrichten (Berlin), 25: 63. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype m, &#8220;Algerien&#8221; [Algeria], ZMHB (Berlin).-Combination <em>Osmia (Chalcosmia) lobata<\/em> Friese in Friese (1911b: 111). Combination <em>Osmia (Diceratosmia) lobata<\/em> Friese in Tkalcu (1969a: 333). Combination <em>Osmia (Caerulosmia) lobata<\/em> Friese in Zanden (1988b: 123).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-<\/strong><strong>Northern Africa:<\/strong> <span style=\"text-decoration: underline\">DZ<\/span>, MA.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Ducke (1900); Zanden (1991a).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Unknown.<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) mongolica <\/em><\/strong><span style=\"font-weight: normal\">Morawitz, 1880<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1880 <em>Osmia <strong>mongolica<\/strong><\/em> Morawitz, Bulletin de l&#8217;Acad\u00e9mie Imp\u00e9riale des Sciences de St.-P\u00e9tersbourg, 26: 382. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype f, by designation of Tkalcu (1995: 137), &#8220;s\u00fcd\u00f6stliche Mongolei&#8221; [Mongolia], ZIN (St. Petersburg).-Combination <em>Osmia (Chalcosmia) mongolica<\/em> Morawitz in Zanden (1988b: 125). Combination <em>Osmia (Helicosmia) mongolica<\/em> Morawitz in Wu (2006: 62).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1930 <em>Osmia<\/em> <em>jacoti<strong> lilacina<\/strong><\/em> Cockerell, The Entomologist (London), 63: 83. <span style=\"text-decoration: underline\">Type material<\/span>: f(f), &#8220;Tsinan, near the Shantung Christian University&#8221; [China].-Synonymy in Zanden (1988b: 125), but see Tkalcu (1995: 137).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Asia: <\/strong><span style=\"text-decoration: underline\">MGL<\/span>, RC(NO)<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Ducke (1900); Tkalcu (1995: 137-138); Wu (2006).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Unknown.<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) moreensis <\/em><\/strong><span style=\"font-weight: normal\">Zanden, 1984<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1984 <em>Osmia (Diceratosmia) <strong>moreensis<\/strong><\/em> Zanden, Reichenbachia (Dresden), 22: 184. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype m, &#8220;Rodini, Rhodos&#8221; [Greece], RMNH (Leiden); paratypes mm, ff.-Combination <em>Osmia (Caerulosmia) moreensis<\/em> Zanden in Zanden (1988b: 123).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe:<\/strong> <span style=\"text-decoration: underline\">GR<\/span>, GR(Aegean Islands, Cr). <strong>Southwestern Asia:<\/strong> IL, RL, SYR, TR.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Warncke (1992b); Zanden (1991a).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<strong><br \/>\n<\/strong><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Possibly oligoletic on Fabaceae; all five pollen samples available so far (from 5 different localities) consist of pollen of Fabaceae (e.g. <em>Trifolium<\/em>) (A. M\u00fcller, unpublished).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) muelleri <\/em><\/strong><span style=\"font-weight: normal\">Rameshkumar &amp; Sardar, 2024<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">2024 <em>Osmia (Pyrosmia) <strong>muelleri<\/strong><\/em> Rameshkumar &amp; Sardar in Sardar et al. <span lang=\"EN-US\">(2014), Zootaxa, 5551: 188<\/span>. <span style=\"text-decoration: underline\">Type material<\/span>: <span lang=\"EN-US\">Holotype m, &#8220;Uttarakhand, Dehradun, Forest Research Institute&#8221; [India], National Zoological Collection (Kolkata); paratypes ff, mm.<\/span><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-<\/strong><span lang=\"EN-US\"><strong>Southwestern Asia:<\/strong><i> <\/i>PAK. <strong>Extralimital:<i> <\/i><\/strong>Oriental (<u>India<\/u>).<\/span><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong><span lang=\"EN-US\">No supplementary or more detailed morphological description known.<\/span><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<strong><br \/>\n<\/strong><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Unknown.<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) oramara <\/em><\/strong><span style=\"font-weight: normal\">Warncke, 1992<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1992 <em>Osmia (Pyrosmia) <strong>oramara<\/strong><\/em> Warncke, Linzer Biologische Beitr\u00e4ge, 24: 908. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Oramar in 1500 m\/Hakkari&#8221; [Turkey], OLML (Linz); paratypes ff.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Southwestern Asia:<\/strong> IL, IR, RL, <span style=\"text-decoration: underline\">TR<\/span>.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known. <strong>Note: <\/strong>Male unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: fissures and cavities between flat stones. Brood cells were found immediately below the soil surface down to a depth of 5cm. Old brood cells are repaired and reused. Up to 100 females were observed to nest closely together. <strong>Nesting material: <\/strong>Cells (8-9mm long) are entirely constructed from leaf pulp, without admixture of sand or small pebbles. They are not built freely between the stones, but are attached to each other or to the substrate. As the substrate also gets covered with chewed leaves, the cell chamber is completely surrounded by leaf pulp. (C. Sedivy, C. Praz and A. M\u00fcller, unpublished).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Flower preferences: <\/strong>Possibly oligolectic on Fabaceae; 14 pollen samples from 4 different localities and three brood cells from the same locality exclusively consist of pollen of <em>Astragalus<\/em> (A. M\u00fcller, unpublished).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) polkruga <\/em><\/strong><span style=\"font-weight: normal\">Warncke, 1992<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1992 <em>Osmia (Pyrosmia) <strong>polkruga<\/strong><\/em> Warncke, Linzer Biologische Beitr\u00e4ge, 24: 915. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;80 km NO Aqaba an der Strasse nach Amman&#8221; [Jordan], OLML (Linz).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Southwestern Asia: <\/strong><span style=\"text-decoration: underline\">JOR<\/span><em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known. <strong>Note: <\/strong>Male unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Flower preferences: <\/strong>Unknown.<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) punica <\/em><\/strong><span style=\"font-weight: normal\">P\u00e9rez, 1895 <\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1895 <em>Osmia <strong>punica<\/strong><\/em> P\u00e9rez, Esp\u00e8ces nouvelles de mellif\u00e8res de Barbarie, p. 11. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype m, by designation of Zanden (1991a: 61), &#8220;Constantine&#8221; [Algeria], MNHN (Paris); paralectotypes mm.-Combination <em>Osmia (Chalcosmia) punica<\/em> P\u00e9rez in Friese (1911b: 113). Combination <em>Osmia (Diceratosmia) punica<\/em> P\u00e9rez in Tkalcu (1969a: 333). Combination <em>Osmia (Caerulosmia) punica<\/em> P\u00e9rez in Zanden (1988b: 124).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Africa: <\/strong><span style=\"text-decoration: underline\">DZ<\/span>, ET, LAR, MA, TN.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Ducke (1900); Zanden (1991a).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Polylectic with a preference for Fabaceae; additional pollen sources include Boraginaceae (e.g. <em>Echium<\/em>) and probably Antirrhineae (A. M\u00fcller, unpublished, based on 14 pollen samples from 11 different localities).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>irherma <\/strong><\/em><span style=\"font-weight: normal\">Warncke, 1992<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1992 <em>Osmia (Pyrosmia) punica <strong>irherma<\/strong><\/em> Warncke, Linzer Biologische Beitr\u00e4ge, 24: 905. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;50 km OSO Marrakesch&#8221; [Morocco], OLML (Linz); paratypes ff.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Africa: <\/strong><span style=\"text-decoration: underline\">MA<\/span><em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known. <strong>Note: <\/strong>Male unknown.<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>punica <\/strong><\/em><span style=\"font-weight: normal\">P\u00e9rez, 1896<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1896 Automatically established nominotypical subspecific taxon (original description and type specimen are identical with those of the species rank taxon of the same name above).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Africa: <\/strong><span style=\"text-decoration: underline\">DZ<\/span>, ET, LAR, MA.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known.<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) saxicola <\/em><\/strong><span style=\"font-weight: normal\">Ducke, 1899<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1899 <em>Osmia <strong>saxicola<\/strong><\/em> Ducke, Entomologische Nachrichten (Berlin), 25: 215. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype m, by designation by Peters (Zanden, 1991a: 63), &#8220;bei Triest&#8221; [Italy], ZMHB (Berlin); paralectotype m. <span style=\"text-decoration: underline\">Type species<\/span> of <em>Viridosmia<\/em> Warncke.-Combination <em>Osmia (Chalcosmia) saxicola<\/em> Ducke in Friese (1911b: 112). Combination <em>Osmia (Diceratosmia) saxicola<\/em> Ducke in Tkalcu (1969a: 333). Combination <em>Osmia (Caerulosmia) saxicola <\/em>Ducke in Zanden (1988b: 124).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1934 <em>Osmia<\/em> <strong><em>caelestina<\/em><\/strong> Benoist, Bulletin de la Soci\u00e9t\u00e9 Entomologique de France, 39: 160. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Asie mineure: Jaffa&#8221; [Israel], MNHN (Paris).-Combination <em>Osmia (Diceratosmia) caelestina<\/em> Benoist in Zanden (1985: 66). Combination <em>Osmia (Caerulosmia) caelestina<\/em> Benoist in Zanden (1988b: 123).-Synonymy in Warncke (1992b: 907).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1937 <em>Osmia (Chalcosmia)<\/em> <strong><em>cypricola<\/em><\/strong> Mavromoustakis, The Annals and Magazine of Natural History (London), 20: 520. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Pera Pedi&#8221; [Cyprus], DAAN (Nicosia); paratypes mm, ff.-Combination <em>Osmia (Diceratosmia) cypricola <\/em>Mavromoustakis in Tkalcu (1969a: 333). Combination <em>Osmia (Caerulosmia) cypricola<\/em> Mavromoustakis in Zanden (1988b: 123).-Synonymy in Warncke (1992b: 907).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1956 <em>Osmia (Chalcosmia)<\/em> <strong><em>posti<\/em><\/strong> Mavromoustakis, The Annals and Magazine of Natural History (London), ser. 12, 9: 856. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Jebel el Knaiss\u00e9, 1700 m&#8221; [Lebanon], DAAN (Nicosia); paratype m.-Combination <em>Osmia (Diceratosmia) posti<\/em> Mavromoustakis in Tkalcu (1969a: 333). Combination <em>Osmia (Caerulosmia) posti<\/em> Mavromoustakis in Zanden (1988b: 124).-Synonymy in Warncke (1992b: 907).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>ARM, E, F, GR, GR(Aegean Islands, Cr),HR, <span style=\"text-decoration: underline\">I<\/span>, SCG, RUS(SR)<em>. <\/em><strong>Northern Asia:<\/strong> TJ. <strong>Southwestern Asia: <\/strong>CY, IL, IR, <span style=\"color: #ff6600\"><span style=\"color: #000000\">IRQ,<\/span><\/span> JOR, RL, SYR, TR<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Ducke (1900); Mavromoustakis (1956a: 857); Warncke (1992b); Zanden (1985: 66, 1991a).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: two nests attached to the undersurface of a stone with 3 and 5 cells. <strong>Nesting material:<\/strong> Cells are entirely constructed with very small pebbles cemented together with chewed leaves. The small orifice between the stone and the soil was observed to be sealed with the same material. (Mavromoustakis, 1948c)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Polylectic with a probable preference for Fabaceae (e.g. <em>Lotus<\/em>); additional pollen sources include Brassicaceae, <em>Allium<\/em> (Alliaceae) and an unknown pollen host (A. M\u00fcller, unpublished, based on 14 pollen samples from 10 different localities).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>riemanni <\/strong><\/em><span style=\"font-weight: normal\">Warncke, 1992<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1992 <em>Osmia saxicola <strong>riemanni<\/strong><\/em> Warncke, Linzer Biologische Beitr\u00e4ge, 24: 908. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Malia\/32 km O Heraklion, Kreta&#8221; [Greece: Crete], OLML (Linz); paratypes ff.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong><span style=\"text-decoration: underline\">GR(Cr)<\/span><em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known. <strong>Note: <\/strong>Male unknown.<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>saxicola <\/strong><\/em><span style=\"font-weight: normal\">Ducke, 1899<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1899 Automatically established nominotypical subspecific taxon (original description and type specimen are identical with those of the species rank taxon of the same name above).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>ARM, E, F, GR, GR(Aegean Islands), HR, <span style=\"text-decoration: underline\">I<\/span>, SCG, RUS(SR)<em>. <\/em><strong>Northern Asia:<\/strong> TJ. <strong>Southwestern Asia: <\/strong>CY, IL, IRQ, JOR, RL, TR<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known.<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) submicans <\/em><\/strong><span style=\"font-weight: normal\">Morawitz, 1870<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1870 <em>Osmia <strong>submicans<\/strong><\/em> Morawitz, Horae Societatis Entomologicae Rossicae (St. Petersburg), 7: 314. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype f, by designation of Zanden (1991a: 64), &#8220;Tauria&#8221; [Ukraine], ZIN (St. Petersburg); paralectotype f.-Combination <em>Osmia (Chalcosmia) submicans<\/em> Morawitz in Friese (1911b: 113). Combination <em>Osmia (Diceratosmia) submicans<\/em> Morawitz in Tkalcu (1969a: 333). Combination <em>Osmia (Caerulosmia) submicans<\/em> Morawitz in Zanden (1988b: 124).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1886 <em>Osmia (Chalcosmia)<\/em> <strong><em>Giraudi<\/em><\/strong> Schmiedeknecht, Apidae Europaeae, vol. 2, p. 117 [983]. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype f, by designation of Zanden (1991a: 64), &#8220;Mallorca, Miramar&#8221; [Spain: Balearic Islands], ZMHB (Berlin).-Synonymy in Ferton (1908: 539).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe:<\/strong> AL, CH, CZ, D, E, E(Ba), F, F(Co), FL, GR, GR(Aegean Islands, Cr), HR, I, I(Sa), I(Si), M, MK, NL, P, RO, RUS(SR), SCG, SK, SLO, <span style=\"text-decoration: underline\">UA<\/span>(Crimea) <strong>Northern Africa:<\/strong> DZ, E(Ca), ET, LAR, MA, TN. <strong>Southwestern Asia:<\/strong> CY, IL, IRQ, JOR, RL, SYR, TR.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Amiet et al. (2004); Banaszak and Romasenko (2001); Benoist (1931); Ducke (1900); Medvedeva (1978); M\u00f3cz\u00e1r (1958); Scheuchl (1996); Schmiedeknecht (1885-1886); Warncke (1992b); Zanden (1991a).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: insect burrows in dead wood and drilled borings in wooden blocks; hollow stems (e.g. <em>Phragmites, Rubus<\/em>); cavities and small holes in rocks and walls; abandoned cells in exposed nests of <em>Megachile (Chalicodoma) parietina<\/em>. <strong>Nesting material:<\/strong> Cell partitions are made of chewed leaves. The nest plug is made of leaf pulp alone or of a mixture of small pebbles (mainly on the inner side of the plug) and leaf pulp. Occasionally, the nest plug consists of alternating layers of leaf pulp and tiny pebbles or mud. In larger cavities, the cells are sometimes arranged side by side with the cell walls partially or entirely built of leaf pulp. (Banaszak and Romasenko, 2001; Benoist, 1931; Ferton, 1908; Gogala, 1999; Krombein, 1969; Maneval, 1939; Vicens, Bosch and Blas, 1993; Wafa and El-Berry, 1971; Westrich, 1989)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Polylectic with a strong preference for Fabaceae; additional pollen sources include Cistaceae, <em>Reseda<\/em> (Resedaceae), <em>Echium<\/em> (Boraginaceae), Lamiaceae (Lamioideae, Nepetoideae), Crassulaceae, Campanulaceae and Antirrhineae (Amiet et al., 2004; Westrich, 1989; A. M\u00fcller, unpublished, based on 45 pollen samples from 37 different localities).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>canaria <\/strong><\/em><span style=\"font-weight: normal\">Mavromoustakis, 1957<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1957 <em>Osmia submicans <strong>canaria<\/strong><\/em> Mavromoustakis, Societas Scientiarum Fennica, Commentationes Biologicae (Helsinki), 16: 1. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Gran Canaria, Arucas&#8221; [Spain: Canary Islands], FMNH (Helsinki).-Combination <em>Osmia (Diceratosmia) canaria<\/em> Mavromoustakis in Tkalcu (1977c: 97). Combination <em>Osmia (Pyrosmia) submicans canaria<\/em> Mavromoustakis in Peters (1975: 50) and in Warncke (1992b: 911).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Africa: <\/strong><span style=\"text-decoration: underline\">E(Ca)<\/span><em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Peters (1975: 50, 54-55); Tkalcu (1977c: 97; 1993b: 816-820); Zanden (1991a, 1996b: 892).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>columbina <\/strong><\/em><span style=\"font-weight: normal\">Zanden, 1996<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1996 <em>Osmia (Pyrosmia) submicans <strong>columbina<\/strong><\/em> Zanden, Linzer Biologische Beitr\u00e4ge, 28: 891. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Puntagorde, Isla La Palma, Cumbercito&#8221; [Spain: Canary Islands], RMNH (Leiden); paratypes mm, f.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Africa: <\/strong><span style=\"text-decoration: underline\">E(Ca)<\/span><em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known.<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>hebraea <\/strong><\/em><span style=\"font-weight: normal\">Benoist, 1934<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1934 <em>Osmia <strong>hebraea<\/strong><\/em> Benoist, Bulletin de la Soci\u00e9t\u00e9 Entomologique de France, 39: 160. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Asie mineure: J\u00e9rusalem&#8221; [Israel], MNHN (Paris).-Combination <em>Osmia (Diceratosmia) hebraea<\/em> Benoist in Tkalcu (1969a: 333). Combination <em>Osmia submicans hebraea<\/em> Benoist in Zanden (1988b: 124).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1969 <em>Osmia<\/em> <strong><em>aurantiaca<\/em><\/strong> Stanek, Annalen des Naturhistorischen Museums in Wien, 73: 379. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype m, &#8220;El Doqqi&#8221; [Egypt], USNM (Washington); paratypes mm.-Combination <em>Osmia (Diceratosmia) submicans aurantiaca<\/em> Stanek in Tkalcu (1977c: 94).-Synonymy in Zanden (1985: 67).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>I(Si), M<em>. <\/em><strong>Northern Africa: <\/strong>DZ, ET, LAR, MA, TN<em>. <\/em><strong>Southwestern Asia: <\/strong>CY, <span style=\"text-decoration: underline\">IL<\/span>, IRQ, JOR, RL, SYR, TR<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Tkalcu (1977c: 94-97); Zanden (1991a, 1996b: 892).<\/p>\n<h3 style=\"text-align: left\"><strong><span style=\"font-weight: normal\"><em>&#8211; <\/em>subspecies<em> <strong>lanzarotae <\/strong><\/em>Warncke, 1992<\/span><\/strong><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1992 <em>Osmia (Pyrosmia) submicans <strong>lanzarotae<\/strong><\/em> Warncke, Linzer Biologische Beitr\u00e4ge, 24: 911. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Penas del Chache, Lanzarote&#8221; [Spain: Canary Islands], OLML (Linz); paratypes ff.<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1993 <em>Osmia (Caerulosmia)<\/em> <strong><em>lanzarotensis<\/em><\/strong> Tkalcu, Ver\u00f6ffentlichungen des \u00dcbersee-Museum Bremen (Naturwissenschaften), 12: 816. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Lanzarote 300 m La Florida 10 km NW Arrecife&#8221; [Spain: Canary Islands], MICN (Santa Cruz de Tenerife); paratypes mm, ff.-New synonymy based on the original description.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Africa: <\/strong><span style=\"text-decoration: underline\">E(Ca)<\/span><em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Zanden (1996b: 892).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>submicans <\/strong><\/em><span style=\"font-weight: normal\">Morawitz, 1870<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1870 Automatically established nominotypical subspecific taxon (original description and type specimen are identical with those of the species rank taxon of the same name above).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>CH, CZ, D, E, E(Ba), F, F(Co), FL, GR, GR(Aegean Islands, Cr), HR, I, I(Sa), MK, NL, P, RO, RUS(SR), SCG, SK, SLO, <span style=\"text-decoration: underline\">UA<\/span>(Crimea).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Mavromoustakis (1957a: 2); Tkalcu (1977c: 97, 1993b: 816-820); Zanden (1996b: 892).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) versicolor <\/em><\/strong><span style=\"font-weight: normal\">Latreille, 1811<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1811 <em>Osmia <strong>versicolor<\/strong><\/em> Latreille, Encyclop\u00e9die M\u00e9thodique, Histoire Naturelle, Insectes, vol. 8, p. 586. <span style=\"text-decoration: underline\">Type material<\/span>: Neotype f, by designation of Zanden (1984c: 182), &#8220;Cattolica&#8221; [Italy], RMNH (Leiden), in original description &#8220;pr\u00e8s de Marseille&#8221; [France], see Warncke (1992b: 915).-Combination <em>Osmia (Chalcosmia) versicolor<\/em> Latreille in Schmiedeknecht (1885: 20 [886]). Combination <em>Osmia (Diceratosmia) versicolor<\/em> Latreille in Tkalcu (1969a: 333). Combination <em>Osmia (Caerulosmia) versicolor<\/em> Latreille in Zanden (1988b: 124).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>AL, ARM, AZ, BG, D, E, E(Ba), F, F(Co), GE, GR, GR(Aegean Islands, Cr), H, HR, <span style=\"text-decoration: underline\">I<\/span>, I(Sa), I(Si), MK, P, RUS(SR), SCG, SLO, UA; <del>UA(Crimea)<\/del>, <span style=\"text-decoration: line-through\">CH<\/span><em>. <\/em><strong>Northern Africa: <\/strong>DZ, LAR, MA, TN<em>. <\/em><strong>Southwestern Asia: <\/strong>IL, IRQ, JOR, RL, SYR, TR<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Amiet et al. (2004); Banaszak and Romasenko (2001); Benoist (1931); Ducke (1900); Medvedeva (1978); M\u00f3cz\u00e1r (1958); Scheuchl (1996); Schmiedeknecht (1885-1886); Warncke (1992b); Zanden (1984c: 184, 1991a).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: empty snail shells of small to medium size (e.g. <em>Bulimus, Cochlicella, Helix, Rumina, Sphincterophila<\/em>) with usually several cells (up to 7) per shell. <strong>Nesting material: <\/strong>Cell partitions and nest plug are made of chewed leaves without addition of pebbles. The space between the partition of the outermost cell and the nest plug is densely filled with small pebbles, earth crumbs or pieces of dry blades. The closed shells are neither moved nor covered with patches of leaf pulp. (Banaszak and Romasenko, 2001; Benoist, 1931; Ferton, 1890, 1901a; Moreno-Rueda et al., 2008; Westrich, 1989; G. Le Goff, personal observation; A. M\u00fcller, personal observation)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Polylectic with a strong preference for Fabaceae; additional pollen sources include Lamiaceae, <em>Echium<\/em> (Boraginaceae), Brassicaceae, Cistaceae, Crassulaceae, Campanulaceae and Asteraceae (Asteroideae, Cichorioideae) (Amiet et al., 2004; Westrich, 1989; A. M\u00fcller, unpublished, based on 44 pollen samples from 37 different localities).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies<\/span><em> <strong>corrusca <\/strong><\/em><span style=\"font-weight: normal\">Erichson, 1835<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1835 <em>Osmia <strong>corrusca<\/strong><\/em> Erichson, in: Waltl, Reise durch Tirol, Oberitalien und Piemont nach dem s\u00fcdlichen Spanien (Passau), vol. 2, p. 107. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype f, by designation of Zanden (1991a: 67), &#8220;Andalusien&#8221; [Spain], ZMHB (Berlin).-Combination <em>Osmia (Caerulosmia) versicolor corrusca<\/em> Erichson in Zanden (1988b: 124).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1857 <em>Megachile<strong> laeta<\/strong><\/em> Gistel, Vacuna, 2: 564. NOMEN PRAEOCCUPATUM (not <em>Megachile laeta <\/em>Smith, 1853). <span style=\"text-decoration: underline\">Type material<\/span>: f(f), &#8220;Algarbia&#8221; [Portugal].-Synonymy in Warncke (1992b: 916).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1917 <em>Megachile<strong> algarbiensis<\/strong><\/em> Strand, Archiv f\u00fcr Naturgeschichte, 82: 98. NOMEN NOVUM with same type specimen for preoccupied <em>Megachile laeta<\/em> Gistel, 1857 [not <em>Megachile laeta <\/em>Smith, 1853].<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe:<\/strong> <span style=\"text-decoration: underline\">E<\/span>, E(Ba), F(Co), P; <span style=\"text-decoration: line-through\">I(Si)<\/span>. <strong>Northern Africa:<\/strong> DZ, LAR, MA, TN.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Warncke (1992b); Zanden (1984c: 183, 1991a).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>versicolor <\/strong><\/em><span style=\"font-weight: normal\">Latreille, 1811<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1811 Automatically established nominotypical subspecific taxon (original description and type specimen are identical with those of the species rank taxon of the same name above).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>AL, ARM, AZ, BG, D, E, F, GE, GR, GR(Aegean Islands, Cr), H, HR, <span style=\"text-decoration: underline\">I<\/span>, I(Sa), I(Si), MK, P, RUS, SCG, SLO, UA; <del>UA(Crimea)<\/del>.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Warncke (1992b).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>viricephalica <\/strong><\/em><span style=\"font-weight: normal\">Warncke, 1992<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1992 <em>Osmia (Pyrosmia) versicolor <strong>viricephalica<\/strong><\/em> Warncke, Linzer Biologische Beitr\u00e4ge, 24: 916. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Efes\/Aydin&#8221; [Turkey], OLML (Linz); paratypes mm, ff.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Southwestern Asia:<\/strong> IL, IRQ, JOR, RL, SYR, <span style=\"text-decoration: underline\">TR<\/span>.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known.<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) viridana <\/em><\/strong><span style=\"font-weight: normal\">Morawitz, 1873<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1873 <em>Osmia <strong>viridana<\/strong><\/em> Morawitz, Horae Societatis Entomologicae Rossicae (St. Petersburg), 10: 150. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype f, by designation of Zanden (1984c: 183), &#8220;Derbent&#8221; [Russia], ZIN (St. Petersburg); paralectotypes mm.-Combination <em>Osmia (Chalcosmia) viridana<\/em> Morawitz in Schmiedeknecht (1885: 20 [886]). Combination <em>Osmia (Diceratosmia) viridana<\/em> Morawitz in Tkalcu (1969a: 333). Combination <em>Osmia (Caerulosmia) viridana<\/em> Morawitz in Zanden (1988b: 124).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1896 <em>Osmia<\/em> <strong><em>angulata<\/em><\/strong> P\u00e9rez, Esp\u00e8ces nouvelles de mellif\u00e8res de Barbarie, p. 11. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype m, see Warncke (1992b: 913), &#8220;Marseille&#8221; [France], MNHN (Paris).-Combination <em>Osmia (Diceratosmia) angulata<\/em> P\u00e9rez in Tkalcu (1969a: 333).-Synonymy in Zanden (1991a: 68).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1984 <em>Osmia (Diceratosmia) viridana<\/em> <strong><em>mulleolus<\/em><\/strong> Zanden, Reichenbachia (Dresden), 22: 184. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Kavarna&#8221; [Bulgaria], RMNH (Leiden); paratypes mm, ff.-Combination <em>Osmia (Caerulosmia) viridana mulleolus<\/em> Zanden in Zanden (1988b: 124).-Synonymy in Warncke (1992b: 913).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe:<\/strong> ARM, AZ, BG, CH, D, E, F, GR, GR(Aegean Islands, Cr), HR, I, MK, <u>RUS<\/u>(SR), SLO, UA(Crimea); Caucasus. <strong>Northern Africa:<\/strong> DZ, TN. <strong>Northern Asia:<\/strong> KS, KZ, TJ, TM, UZ. <strong>Southwestern Asia: <\/strong>CY, IL, IR, IRQ, JOR, RL, SYR, TR.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Amiet et al. (2004); Banaszak and Romasenko (2001); Benoist (1931); Ducke (1900); Medvedeva (1978); M\u00f3cz\u00e1r (1958); Scheuchl (1996); Schmiedeknecht (1885-1886); Warncke (1992b); Zanden (1984c: 184, 1991a).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: empty snail shells of small to medium size (e.g. <em>Eobania, Helicella, Zebrina<\/em>) with usually several cells per shell; holes in rocks; abandoned cells in exposed nests of <em>Megachile (Chalicodoma) lefeburei <\/em>with up to 7 cells per <em>Megachile<\/em> cell. <strong>Nesting material: <\/strong>Cell partitions and nest plug are made of chewed leaves mixed with small pebbles. The space between the partition of the outermost cell and the nest plug is loosely filled with small pebbles, earth crumbs or pieces of dry blades. (Amiet et al., 2004; Banaszak and Romasenko, 2001; Benoist, 1931; Ferton, 1901a; Mavromoustakis, 1939b, 1948c, 1957b; A. M\u00fcller, personal observation)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Polylectic with a strong preference for Fabaceae; additional pollen sources include Lamiaceae, <em>Echium<\/em> (Boraginaceae), Brassicaceae, Crassulaceae and monocots (Amiet et al., 2004; Westrich, 1989; A. M\u00fcller, unpublished, based on 49 pollen samples from 37 different localities).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>nicosiana <\/strong><\/em><span style=\"font-weight: normal\">Mavromoustakis, 1939<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1939 <em>Osmia (Chalcosmia) <strong>nicosiana<\/strong><\/em> Mavromoustakis, The Annals and Magazine of Natural History (London), ser. 11, 3: 155. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Ayia Phyla&#8221; [Cyprus], DAAN (Nicosia); paratypes mm, ff.-Combination <em>Osmia (Caerulosmia) nicosiana<\/em> Mavromoustakis in Zanden (1988b: 124). Combination <em>Osmia (Pyrosmia) viridana nicosiana<\/em> Mavromoustakis in Warncke (1992b: 913).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Southwestern Asia: <\/strong><span style=\"text-decoration: underline\">CY<\/span>.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Warncke (1992b); Zanden (1991a).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>rufispina <\/strong><\/em><span style=\"font-weight: normal\">Morawitz, 1875<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1875 <em>Osmia <strong>rufispina<\/strong><\/em> Morawitz, Travel to Turkestan by A. P. Fedtschenko, p. 83. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;in valle Sarafschan&#8221; [Tajikistan], ZMUM (Moscow).-Combination <em>Osmia (Caerulosmia) rufispina<\/em> Morawitz in Zanden (1988b: 124). Combination <em>Osmia (Pyrosmia) viridana rufispina<\/em> Morawitz in Warncke (1992b: 913).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Asia: <\/strong>KS, KZ, <span style=\"text-decoration: underline\">TJ<\/span>, TM, UZ.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Warncke (1992b); Zanden (1991a). <strong>Note: <\/strong>Male unknown.<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>viridana <\/strong><\/em><span style=\"font-weight: normal\">Morawitz, 1873<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1873 Automatically established nominotypical subspecific taxon (original description and type specimen are identical with those of the species rank taxon of the same name above).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>ARM, AZ, BG, CH, D, E, F, GR, GR(Aegean Islands, Cr), HR, I, MK, <u>RUS<\/u>(SR), SLO, UA(Crimea); Caucasus. <strong>Northern Africa:<\/strong> DZ, TN. <strong>Southwestern Asia:<\/strong> IL, IR, IRQ, JOR, RL, SYR, TR.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Zanden (1991a).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) zarzisa <\/em><\/strong><span style=\"font-weight: normal\">Warncke, 1992<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1992 <em>Osmia (Pyrosmia) <strong>zarzisa<\/strong><\/em> Warncke, Linzer Biologische Beitr\u00e4ge, 24: 913. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype m, &#8220;30 km S Zarzis&#8221; [Tunisia], OLML (Linz); paratypes mm, f.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Africa:<\/strong> <span style=\"text-decoration: underline\">TN<\/span>.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Unknown.<\/p>\n<h3 style=\"text-align: left\"><span style=\"text-decoration: underline\"><em>cephalotes<\/em> species group<\/span><\/h3>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) cephalotes <\/em><\/strong><span style=\"font-weight: normal\">Morawitz, 1870<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1870 <em>Osmia <strong>cephalotes<\/strong><\/em> Morawitz, Horae Societatis Entomologicae Rossicae (St. Petersburg), 7: 315. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Gallia merid., in der Umgebung von Nizza&#8221; [France], ZIN (St. Petersburg).-Combination <em>Osmia (Chalcosmia) cephalotes<\/em> Morawitz in Schmiedeknecht (1885: 20 [886]). Combination <em>Osmia (Caerulosmia) cephalotes<\/em> Morawitz in Zanden (1988b: 123).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1879 <em>Osmia<\/em> <strong><em>bacillus<\/em><\/strong> P\u00e9rez, Actes de la Soci\u00e9t\u00e9 Linn\u00e9enne de Bordeaux, 33: 203. <span style=\"text-decoration: underline\">Type material<\/span>: f(f), &#8220;Marseille&#8221; [France], MNHN (Paris).-Synonymy with <em>Osmia longiceps<\/em> Morawitz in Dalla Torre (1896: 401).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1899 <em>Osmia<\/em> <strong><em>pulsata<\/em><\/strong> Buysson, Annales de la Soci\u00e9t\u00e9 Entomologique de France, 68: 668. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype m, &#8220;Tunis&#8221; [Tunisia], MNHN (Paris).-Synonymy in Zanden (1986: 73).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe:<\/strong> ARM, AZ, BG, E, <span style=\"text-decoration: underline\">F<\/span>, GE, GR, GR(Aegean Islands), H, HR, I, I(Si), P, RO, RUS(SR), SCG, SLO, UA, UA(Crimea). <strong>Northern Africa:<\/strong> DZ, LAR, MA, TN. <strong>Northern Asia: <\/strong>TM. <strong>Southwestern Asia: <\/strong>CY, IL, IR, <span style=\"color: #ff6600\"><span style=\"color: #000000\">IRQ, <\/span><\/span>JOR, RL, SYR, TR.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Banaszak and Romasenko (2001); Benoist (1931); Ducke (1897: 40); M\u00f3cz\u00e1r (1958); Warncke (1992b); Zanden (1991a).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: insect burrows in dead wood. <strong>Nesting material:<\/strong> Cell partitions and nest plug are made of chewed leaves. Females were also observed to collect pebbles and mud on the ground, indicating that this species occasionally uses a mixture of leaf pulp and pebbles as nest building material as many other <em>Pyrosmia<\/em> species do or that the pebbles are used to barricade the nest entrance, a behaviour that is known from other <em>Pyrosmia<\/em> species as well. (Benoist, 1931; Ferton, 1895; G. Le Goff, personal observation; A. M\u00fcller, personal observation)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Polylectic with a preference for Fabaceae; additional pollen sources include <em>Echium<\/em> (Boraginaceae), Lamiaceae (Lamioideae, Nepetoideae), monocots and Cistaceae (A. M\u00fcller, unpublished, based on 26 pollen samples from 25 different localities and on field observations). Flower records: <em>Salvia fruticosa<\/em> (label record).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>cephalotes <\/strong><\/em><span style=\"font-weight: normal\">Morawitz, 1870<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1870 Automatically established nominotypical subspecific taxon (original description and type specimen are identical with those of the species rank taxon of the same name above).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>E, <span style=\"text-decoration: underline\">F<\/span>, I, I(Si), P<em>.<\/em><strong> Northern Africa:<\/strong> DZ, LAR, MA, TN.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Ducke (1900); Schmiedeknecht (1885-1886); Zanden (1991a).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies<\/span><em> <strong>longiceps <\/strong><\/em><span style=\"font-weight: normal\">Morawitz, 1875<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1875 <em>Osmia <strong>longiceps<\/strong><\/em> Morawitz, Horae Societatis Entomologicae Rossicae (St. Petersburg), 12: 40. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype f, by designation of Zanden (1991a: 51), &#8220;Borshom&#8221; [Georgia], ZIN (St. Petersburg).-Combination <em>Osmia (Chalcosmia) longiceps<\/em> Morawitz in Schmiedeknecht (1885: 20 [886]).-Synonymy with <em>Osmia cephalotes<\/em> Morawitz in Alfken (1914: 201).-Combination <em>Osmia (Caerulosmia) cephalotes longiceps<\/em> Morawitz in Zanden (1988b: 123).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe:<\/strong> ARM, AZ, BG, <span style=\"text-decoration: underline\">GE<\/span>, GR, GR(Aegean Islands), H, HR, I, RO, RUS(SR), SCG, SLO, UA, UA(Crimea). <strong>Northern Asia: <\/strong>TM. <strong>Southwestern Asia: <\/strong>CY, IL, IR, <span style=\"color: #ff6600\"><span style=\"color: #000000\">IRQ, <\/span><\/span>JOR, RL, SYR, TR.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Ducke (1900); Schmiedeknecht (1885-1886); Zanden (1991a).<\/p>\n<h3 style=\"text-align: left\"><span style=\"text-decoration: underline\"><em>gallarum<\/em> species group<\/span><\/h3>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) gallarum <\/em><\/strong><span style=\"font-weight: normal\">Spinola, 1808<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1808 <em>Osmia <strong>gallarum<\/strong><\/em> Spinola, Insectorum Liguriae, vol. 2, p. 69. <span style=\"text-decoration: underline\">Type material<\/span>: Neotype f, by designation of Zanden (1984c: 187), &#8220;Venezia-Ciulia&#8221; [Italy], RMNH (Leiden).-<span style=\"text-decoration: underline\">Type species<\/span> of <em>Caerulosmia<\/em> Zanden.-Combination <em>Osmia (Chalcosmia) gallarum<\/em> Spinola in Schmiedeknecht (1885: 20 [886]). Combination <em>Osmia (Diceratosmia) gallarum<\/em> Spinola in Tkalcu (1969a: 333). Combination <em>Osmia (Caerulosmia) gallarum<\/em> Spinola in Zanden (1988b: 123).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1840 <em>Osmia<\/em> <strong><em>ruborum<\/em><\/strong> Dufour &amp; Perris, Annales de la Soci\u00e9t\u00e9 Entomologique de France, 9: 13. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype f, by designation of Zanden (1991a: 56), &#8220;Gallia meridionali-occidentali (Landes)&#8221; [France], MNHN (Paris).-Synonymy in Zanden (1991a: 56).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>A, AL, B, BG, CH, CZ, D, E, F, GR, H, HR, <span style=\"text-decoration: underline\">I<\/span>, L, MK, P, PL, RO, SCG, SK, SLO; <del>UA(Crimea)<\/del><em>. <\/em><strong>Northern Africa: <\/strong>DZ, MA, TN<em>. <\/em><strong>Southwestern Asia: <\/strong>TR<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Amiet et al. (2004); Banaszak and Romasenko (2001); Benoist (1931); Ducke (1900); Mavromoustakis (1956a: 857); Medvedeva (1978); M\u00f3cz\u00e1r (1958); Scheuchl (1996); Schmiedeknecht (1885-1886); Warncke (1992b); Zanden (1984c: 186-191, 1991a).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology-Nesting site: <\/strong>Preexisting cavities: insect burrows in dead wood and drilled borings in wooden blocks; hollow stems (e.g. <em>Rubus, Sambucus<\/em>) and drilled borings in pithy stems; cavities within abandoned galls of <em>Andricus kollari<\/em> with 10-24 cells and of <em>Cynips argentea<\/em> with 10-12 cells. It is unclear whether the females are able to gnaw burrows of their own in the pith of stems or galls. In galls, pith is sometimes removed to accomodate the cells, which are irregularly arranged within the cavity of the galls. <strong>Nesting material: <\/strong>Cell partitions are made of chewed leaves. The nest plug consists of three to four partitions of leaf pulp separated by 0.5-1mm from each other. When nesting in pithy stems, pith is sometimes incorporated into the leaf matrix of partitions and plug. In the subspecies <em>lapidistructor<\/em>, the cell partitions are built of small pebbles cemented together with leaf pulp. (Banaszak and Romasenko, 2001; Benoist, 1931; Ducke, 1900; Enslin, 1925, 1933; Ferton, 1920; Friese, 1911b, 1923; Latreille, 1811; Westrich, 1989)<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Oligolectic on Fabaceae (Amiet et al., 2004; Westrich, 1989; A. M\u00fcller, unpublished, based on 6 pollen samples from 6 different localities and on field observations).<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies <\/span><em><strong>gallarum <\/strong><\/em><span style=\"font-weight: normal\">Spinola, 1808<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1808 Automatically established nominotypical subspecific taxon (original description and type specimen are identical with those of the species rank taxon of the same name above).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>A, AL, B, BG, CH, CZ, D, E, F, GR, H, HR, <span style=\"text-decoration: underline\">I<\/span>, L, MK, P, PL, RO, SCG, SK, SLO; <del>UA(Crimea)<\/del><em>. <\/em><strong>Southwestern Asia: <\/strong>TR<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known.<\/p>\n<h3 style=\"text-align: left\"><em><span style=\"font-weight: normal\">&#8211; <\/span><\/em><span style=\"font-weight: normal\">subspecies<\/span><em> <strong>lapidistructor <\/strong><\/em><span style=\"font-weight: normal\">Ferton, 1920<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1920 <em>Osmia <strong>lapidistructor<\/strong><\/em> Ferton, Annales de la Soci\u00e9t\u00e9 Entomologique de France (Paris), 89: 343. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;La Calle&#8221; [Algeria], MNHN (Paris).-Synonymy with <em>Osmia gallarum<\/em> Spinola in Zanden (1988b: 123).-Combination <em>Osmia gallarum lapidistructor<\/em> Ferton in Warncke (1992b: 904).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Africa: <\/strong><span style=\"text-decoration: underline\">DZ<\/span>, MA, TN<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Warncke (1992b).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) hellados <\/em><\/strong><span style=\"font-weight: normal\">Zanden, 1984<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1984 <em>Osmia (Diceratosmia) <strong>hellados<\/strong><\/em> Zanden, Reichenbachia (Dresden), 22: 189. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype m, &#8220;Palaiokastritsa, Korfu&#8221; [Greece], RMNH (Leiden); paratypes mm, ff.-Combination <em>Osmia (Caerulosmia) hellados<\/em> Zanden in Zanden (1988b: 115). Combination <em>Osmia (Pyrosmia) gallarum hellados<\/em> Zanden in Warncke (1992b: 903). Combination <em>Osmia hellados<\/em> Zanden in Standfuss <em>et al.<\/em> (2003: 302).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe: <\/strong>ARM, AZ, BG, GE, <span style=\"text-decoration: underline\">GR<\/span>, GR(Aegean Islands, Cr), HR, I, I(Si), MK, RUS(SR), UA(Crimea); Yugoslavia<em>. <\/em><strong>Southwestern Asia: <\/strong>CY, IL, JOR, RL, TR<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Warncke (1992b); Zanden (1991a).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Unknown.The only pollen sample available so far consists of pollen of Fabaceae (A. M\u00fcller, unpublished).<\/p>\n<h3 style=\"text-align: left\"><span style=\"text-decoration: underline\"><em>nana<\/em> species group<\/span><\/h3>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) avedata <\/em><\/strong><span style=\"font-weight: normal\">Warncke, 1992<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1992 <em>Osmia (Pyrosmia) <strong>avedata<\/strong><\/em> Warncke, Linzer Biologische Beitr\u00e4ge, 24: 901. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Avedat&#8221; [Israel], OLML (Linz); paratypes m(m), f(f).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Southwestern Asia: <\/strong>JOR, <span style=\"text-decoration: underline\">IL<\/span>.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Unknown.<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) nana <\/em><\/strong><span style=\"font-weight: normal\">Morawitz, 1873<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1873 <em>Osmia <strong>nana <\/strong><\/em>Morawitz, Horae Societatis Entomologicae Rossicae (St. Petersburg), 10: 152. <span style=\"text-decoration: underline\">Type material<\/span>: Lectotype m, by designation of Zanden (1981: 154), &#8220;Derbent&#8221; [Russia], ZIN (St. Petersburg).-Combination <em>Osmia (Chalcosmia) nana<\/em> Morawitz in Friese (1911b: 111). Combination <em>Osmia (Diceratosmia) nana<\/em> Morawitz in Zanden (1981: 153). Combination <em>Osmia (Caerulosmia) nana<\/em> Morawitz in Zanden (1988b: 123).<\/p>\n<p style=\"padding-left: 30px;text-align: left\">1934 <em>Osmia<\/em> <strong><em>tetrodonta<\/em><\/strong> Benoist, Bulletin de la Soci\u00e9t\u00e9 Entomologique de France, 39: 106. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype m, &#8220;Asie mineure: Betmeri&#8221; [Lebanon], MNHN (Paris).-Synonymy in Zanden (1985: 62).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe:<\/strong> AZ, BG, F, GR, GR(Aegean Islands, Cr), HR, I, I(Sa), I(Si), <u>RUS<\/u>(SR); Caucasus. <strong>Northern Africa:<\/strong> DZ?. <strong>Northern Asia:<\/strong> Turkestan. <strong>Southwestern Asia:<\/strong> CY, IL, IR, <span style=\"color: #ff6600\"><span style=\"color: #000000\">IRQ, <\/span><\/span>JOR, RL, SYR, TR<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Ducke (1900); Medvedeva (1978); Stanek (1969a: 39-40); Warncke (1992b); Zanden (1981: 153-156, 1985: 61, 1991a).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Polylectic with a strong preference for Fabaceae; additional pollen sources include Lamiaceae and an unknown pollen host (A. M\u00fcller, unpublished, based on 30 pollen samples from 28 different localities).<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) nigricollis <\/em><\/strong><span style=\"font-weight: normal\">Warncke, 1992<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1992 <em>Osmia (Pyrosmia) <strong>nigricollis<\/strong><\/em> Warncke, Linzer Biologische Beitr\u00e4ge, 24: 900. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Semdinli in 1700 m\/Hakkari&#8221; [Turkey], OLML (Linz); paratypes m(m), f(f).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Southwestern Asia: <\/strong>IR, <span style=\"text-decoration: underline\">TR<\/span><em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Unknown.<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) ramona <\/em><\/strong><span style=\"font-weight: normal\">Warncke, 1992<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1992 <em>Osmia (Pyrosmia) <strong>ramona<\/strong><\/em> Warncke, Linzer Biologische Beitr\u00e4ge, 24: 903. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;Wadi Ramon\/6 km O Mizpe Ramon&#8221; [Israel], OLML (Linz); paratype f.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Southwestern Asia: <\/strong><span style=\"text-decoration: underline\">IL<\/span><em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known. <strong>Note: <\/strong>Male unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Flower preferences: <\/strong>Unknown.<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) tadjika <\/em><\/strong><span style=\"font-weight: normal\">Warncke, 1992 <\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1992 <em>Osmia (Pyrosmia) <strong>tadjika<\/strong><\/em> Warncke, Linzer Biologische Beitr\u00e4ge, 24: 901. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;20 km W Tawil-Dara in 2000 m&#8221; [Tajikistan], OLML (Linz); paratype f.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Asia: <\/strong><span style=\"text-decoration: underline\">TJ<\/span><em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known. <strong>Note: <\/strong>Male unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Flower preferences: <\/strong>Unknown.<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) tawildara <\/em><\/strong><span style=\"font-weight: normal\">Warncke, 1992<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1992 <em>Osmia (Pyrosmia) <strong>tawildara<\/strong><\/em> Warncke, Linzer Biologische Beitr\u00e4ge, 24: 900. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype f, &#8220;20 km W Tawil-Dara in 2000 m&#8221; [Tajikistan], OLML (Linz); paratype f.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Northern Asia: <\/strong><span style=\"text-decoration: underline\">TJ<\/span><em>. <\/em><strong>Southwestern Asia: <\/strong>TR<em>.<\/em><\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known. <strong>Note: <\/strong>Male unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Flower preferences: <\/strong>Unknown.<\/p>\n<h3 style=\"text-align: left\"><strong><em>Osmia (Pyrosmia) teunisseni <\/em><\/strong><span style=\"font-weight: normal\">Zanden, 1981<\/span><\/h3>\n<p style=\"padding-left: 30px;text-align: left\">1981 <em>Osmia (Diceratosmia) <strong>teunisseni<\/strong><\/em> Zanden, Reichenbachia (Dresden), 19: 156. <span style=\"text-decoration: underline\">Type material<\/span>: Holotype m, &#8220;Samos, Manolates&#8221; [Greece], RMNH (Leiden); paratypes mm, ff.-Combination <em>Osmia (Caerulosmia) teunisseni<\/em> Zanden in Zanden (1988b: 124).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Distribution-Europe<\/strong>: <span style=\"text-decoration: underline\">GR<\/span>, GR(Aegean Islands), HR, I. <strong>Southwestern Asia:<\/strong> CY, IL, IR, <span style=\"color: #ff6600\"><span style=\"color: #000000\">IRQ, <\/span><\/span>JOR, RL, SYR, TR.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Identification-<\/strong><strong>Keys, Descriptions: <\/strong>Warncke (1992b); Zanden (1991a).<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong>Nesting biology: <\/strong>Unknown.<\/p>\n<p style=\"padding-left: 30px;text-align: left\"><strong> Flower preferences: <\/strong>Polylectic; pollen sources recorded so far include Fabaceae, Lamiaceae and Crassulaceae (A. M\u00fcller, unpublished, based on 5 pollen samples from 5 different localities).<\/p>\n","protected":false},"excerpt":{"rendered":"<p>The subgenus Pyrosmia Tkalcu is confined to the Palaearctic region. It contains 33 described species. Female of Osmia (Pyrosmia) ferruginea Latreille, 1811. Foto N. Vereecken. Species accounts Osmia (Pyrosmia) amathusica Mavromoustakis, 1937 1937 Osmia (Chalcosmia) amathusica Mavromoustakis, Annals and Magazine of Natural History (London), 20: 522. Type material: Holotype f, &#8220;Amathus (ruins)&#8221; [Cyprus], DAAN (Nicosia); [&hellip;]<\/p>\n","protected":false},"author":1092,"featured_media":0,"parent":46,"menu_order":0,"comment_status":"open","ping_status":"closed","template":"","meta":{"footnotes":""},"class_list":["post-162","page","type-page","status-publish","hentry"],"_links":{"self":[{"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/pages\/162","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/pages"}],"about":[{"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/types\/page"}],"author":[{"embeddable":true,"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/users\/1092"}],"replies":[{"embeddable":true,"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/comments?post=162"}],"version-history":[{"count":5,"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/pages\/162\/revisions"}],"predecessor-version":[{"id":7344,"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/pages\/162\/revisions\/7344"}],"up":[{"embeddable":true,"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/pages\/46"}],"wp:attachment":[{"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/media?parent=162"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}