{"id":144,"date":"2009-01-22T00:03:29","date_gmt":"2009-01-21T23:03:29","guid":{"rendered":"https:\/\/wpethzprd.ethz.ch\/osmiini\/?page_id=144"},"modified":"2026-04-12T16:54:46","modified_gmt":"2026-04-12T15:54:46","slug":"hoplosmia","status":"publish","type":"page","link":"https:\/\/blogs.ethz.ch\/osmiini\/palaearctic-species\/osmia\/hoplosmia\/","title":{"rendered":"Subgenus Hoplosmia"},"content":{"rendered":"

The subgenus Hoplosmia<\/em> Thomson is confined to the Palaearctic region. It contains 21 species (M\u00fcller, 2018a).<\/p>\n

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Female of Osmia (Hoplosmia) spinulosa<\/em> (Kirby, 1802) collecting pollen on a flower head of Buphthalmum salicifolium<\/em> (Asteraceae). Foto A. M\u00fcller.<\/span><\/h5>\n

Species accounts<\/h3>\n

spinulosa<\/u><\/em> species group<\/u><\/strong><\/h3>\n

Osmia (Hoplosmia) <\/em>carbo<\/em> (Zanden, 1994)<\/strong><\/h3>\n

1994 Hoplosmia (Paranthocopa) pinguis <\/em>carbo<\/em><\/strong> Zanden, Linzer Biologische Beitr\u00e4ge, 26: 1116. Type material<\/u>: Holotype f, \u201e8 km O Yeroham\u201c [Israel], Schwarz Collection (Ansfelden); paratypes mm, ff.-Osmia carbo<\/em> (Zanden) stat. nov. in M\u00fcller (2018a).<\/p>\n

Distribution-Northern Africa: <\/strong>ET. Southwestern Asia:<\/strong> IL<\/u>, JOR, SYR.<\/p>\n

Identification-Keys, Descriptions: <\/strong>M\u00fcller (2018a).<\/p>\n

Nesting biology-Nesting site: <\/strong>Preexisting cavities: empty snail shells of small to medium size with 1-3 brood cells. Similar to O<\/em>. <\/em>spinulosa<\/em>, the nests are not sealed against their rear end with a basal wall and the outermost cell partition is distinctly more robust than all the other cell partitions and the nest plug, probably acting as a barrier against parasites and predators. In contrast to O<\/em>. <\/em>spinulosa<\/em>, the shells are never turned after they have been sealed. Nesting material: <\/strong>Cell partitions and nest plug are made of chewed leaves. The shell surface is not plastered with patches of leaf pulp. (M\u00fcller, 2018a and references therein)<\/p>\n

Flower preferences:<\/strong> Oligolectic on Asteraceae with a near exclusive preference for Cichorioideae and Asteroideae; Carduoideae are very rarely exploited as well (M\u00fcller, 2018a).<\/p>\n

Osmia (Hoplosmia) <\/em>centaureae<\/em> M\u00fcller, 2018<\/strong><\/h3>\n

2018 Osmia <\/em>(Hoplosmia<\/em>) centaureae <\/em><\/strong>M\u00fcller, Zootaxa, 4415: 303. Type material<\/u>: Holotype m, \u201eWadi Mujib, King\u2019s Highway\u201c [Jordan], ETHZ (Zurich); paratypes mm, ff.<\/p>\n

Distribution-Southwestern Asia:<\/strong> IL, JOR<\/u>.<\/p>\n

Identification-Keys, Descriptions: <\/strong>No supplementary or more detailed morphological description known.<\/p>\n

Nesting biology: <\/strong>Unknown.<\/p>\n

Flower preferences: <\/strong>Oligolectic on Asteraceae with a probably exclusive preference for Carduoideae (M\u00fcller, 2018a).<\/p>\n

Osmia (Hoplosmia) croatica<\/em> Friese, 1893<\/strong><\/h3>\n

1893 Osmia <\/em>croatica<\/em><\/strong> Friese, Entomologische Nachrichten (Berlin), 19: 353. Type material<\/u>: Lectotype m, by designation of Tkalcu (1974a: 123), \u201eTriest\u201c [Italy], ZMHB (Berlin).-Combination Anthocopa (Hoplosmia) croatica<\/em> (Friese) in Tkalcu (1974a: 122). Combination Hoplosmia (Hoplosmia) croatica<\/em> (Friese) in Ungricht et al. (2008: 131).<\/p>\n

Distribution-Europe:<\/strong> ARM, BG, GR, GR(Aegean Islands), HR, I<\/u>, SCG, SLO. Southwestern Asia:<\/strong> TR.<\/p>\n

Identification-Keys, Descriptions: <\/strong>Ducke (1900); Stanek (1969a: 19-22); Tkalcu (1974a); M\u00fcller (2018a).<\/p>\n

Nesting biology-Nesting site: <\/strong>Preexisting cavities: empty snail shells. The only nest recorded so far contained a single brood cell and was hidden below stones. The nest architecture appears to be exactly the same as in Osmia (Hoplosmia) spinulosa. <\/em>Nesting material: <\/strong>Cell partitions and nest plug are made of chewed leaves (e.g. from Scabiosa<\/em>). The shell surface is not plastered with patches of leaf pulp. (A. M\u00fcller, 2018a)<\/p>\n

Flower preferences: <\/strong>Oligolectic on Asteraceae with a probably exclusive preference for Carduoideae (M\u00fcller, 2018a).<\/p>\n

Osmia (Hoplosmia) elegans<\/em> (Tkalcu, 1992)<\/h3>\n

1992 Hoplosmia (Hoplosmia) <\/em>elegans<\/em><\/strong> Tkalcu, Acta Societatis Zoologicae Bohemoslovacae (Praha), 56: 220. Type material<\/u>: Holotype m, \u201eSide\u201c [Turkey], RMNH (Leiden); paratypes m(m), f(f).<\/p>\n

Distribution-Europe:<\/strong> GR(Lesbos). Southwestern Asia:<\/strong> JOR, TR<\/u>.<\/em><\/p>\n

Identification-Keys, Descriptions: <\/strong>M\u00fcller (2018a).<\/p>\n

Nesting biology: <\/strong>Unknown.<\/p>\n

Flower preferences: <\/strong>The only pollen load analyzed so far consisted of pollen of Carduoideae (Asteraceae) (M\u00fcller, 2018a).<\/p>\n

Osmia (Hoplosmia) fallax<\/em> P\u00e9rez, 1895<\/h3>\n

1895 Osmia <\/em>fallax<\/em><\/strong> P\u00e9rez, Esp\u00e8ces nouvelles de mellif\u00e8res de Barbarie, p. 13. Type material<\/u>: Lectotype m, by designation of Tkalcu (1974a: 131), \u201eAndalusia\u201c [Spain], MNHN (Paris). Type species<\/u> of Odonterythrosmia<\/em> Tkalcu.-Combination Anthocopa (Odonterythrosmia) fallax<\/em> (P\u00e9rez) in Tkalcu (1974a: 131). Combination Hoplosmia (Odontanthocopa) fallax<\/em> (P\u00e9rez) in Ungricht et al. (2008: 134).<\/p>\n

Distribution-Europe:<\/strong> E<\/u>. <\/em>Northern Africa:<\/strong> DZ, MA.<\/p>\n

Identification-Keys, Descriptions: <\/strong>Ducke (1900); Tkalcu (1974a); M\u00fcller (2018a).<\/p>\n

Nesting biology-Nesting site: <\/strong>Preexisting cavities: empty snail shells (e.g. Sphincterophila candidissima<\/em>) with 1-4 cells per shell. Nesting material: <\/strong>Unknown. (M\u00fcller, 2018a and references therein). Brood parasites:<\/strong> possibly Stelis odontopyga<\/em> (Hymenoptera, Megachilidae) (Moreno-Rueda et al., 2008; M\u00fcller, 2018a).<\/p>\n

Flower preferences: <\/strong>Polylectic with a strong preference for Cistaceae (e.g. Helianthemum<\/em>), additional pollen sources include Brassicaceae, Asteraceae (Asteroideae, Cichorioideae), Echium<\/em> (Boraginaceae) and Lamiaceae (M\u00fcller, 2018a).<\/p>\n

Osmia (Hoplosmia) picena<\/em> (Tkalcu, 1999)<\/h3>\n

1999 Hoplosmia (Hoplosmia) <\/em>picena<\/em><\/strong> Tkalcu, Bulletin de la Soci\u00e9t\u00e9 Entomologique de Mulhouse, Janvier-Mars: 11. Type material<\/u>: Holotype f, \u201eP.N. Sibillini\u201c [Italy], RMNH (Leiden); paratypes ff.<\/p>\n

Distribution-Europe:<\/strong> I<\/u>.<\/em><\/p>\n

Identification-Keys, Descriptions: <\/strong>M\u00fcller (2018a).<\/p>\n

Nesting biology: <\/strong>Unknown.<\/p>\n

Flower preferences: <\/strong>Oligolectic on Asteraceae with a preference for Carduoideae; species of the Cichorioideae and the Asteroideae are occasionally also exploited as pollen sources (M\u00fcller, 2018a).<\/p>\n

Osmia (Hoplosmia) pinguis<\/em> P\u00e9rez, 1895<\/h3>\n

1895 Osmia <\/em>pinguis<\/em><\/strong> P\u00e9rez, Esp\u00e8ces nouvelles de mellif\u00e8res de Barbarie, p. 12. Type material<\/u>: Lectotype f, by designation of Tkalcu (1974a: 133), \u201eDa Calle\u201c [Algeria], MNHN (Paris). Type species<\/u> of Paranthocopa<\/em> Tkalcu.-Combination Anthocopa (Paranthocopa) pinguis<\/em> (P\u00e9rez) in Tkalcu (1974a: 133). Combination Hoplosmia (Paranthocopa) pinguis<\/em> (P\u00e9rez) in Ungricht et al. (2008: 136).<\/p>\n

1928 Osmia<\/em> indivisa<\/em><\/strong> Benoist, Bulletin de la Soci\u00e9t\u00e9 des Sciences Naturelles du Maroc (Rabat), 8: 214. Type material<\/u>: Holotype m, \u201eMarrakech\u201c [Morocco], MNHN (Paris).-Synonymy in Zanden (1985: 55).<\/p>\n

Distribution- Europe:<\/strong> E. Northern Africa:<\/strong> DZ<\/u>, MA, TN. <\/em><\/p>\n

Identification-Keys, Descriptions: <\/strong>Ducke (1900); Tkalcu (1974a); M\u00fcller (2018a).<\/p>\n

Nesting biology-Nesting site: <\/strong>Preexisting cavities: empty snail shells of small to medium size (e.g. Otala, Rumina<\/em>) with 2-4 brood cells per shell. Similar to O<\/em>. <\/em>spinulosa<\/em>, the nests are not sealed against their rear end with a basal wall and the outermost cell partition is distinctly more robust than all the other cell partitions and the nest plug, probably acting as a barrier against parasites and predators. In contrast to O<\/em>. <\/em>spinulosa<\/em>, the shells are never turned after they have been sealed. Nesting material: <\/strong>Cell partitions and nest plug are made of chewed leaves. The shell surface is not plastered with patches of leaf pulp. (M\u00fcller, 2018a and references therein)<\/p>\n

Flower preferences: <\/strong>Oligolectic on Asteraceae with a near exclusive preference for Cichorioideae and Asteroideae (M\u00fcller, 2018a).<\/p>\n

Osmia (Hoplosmia) spinigera<\/em> Latreille, 1811<\/h3>\n

1811 Osmia <\/em>spinigera<\/em><\/strong> Latreille, Encyclop\u00e9die M\u00e9thodique, Histoire Naturelle, Insectes, vol. 8, p. 584. Type material<\/u>: Syntypes m(m), f(f), \u201e\u00c9gypte\u201c [Egypt], Olivier Collection.-Combination Anthocopa (Hoplosmia) spinigera<\/em> (Latreille) in Tkalcu (1974a: 122). Combination Hoplosmia (Hoplosmia) spinigera<\/em> (Latreille) in Ungricht et al. (2008: 131).<\/p>\n

1869 Osmia<\/em> clavicula<\/em><\/strong> Gerstaecker, Entomologische Zeitung (Stettin), 30: 347. Type material<\/u>: Lectotype m, by designation of Tkalcu (1974a: 122), \u201eNaxos\u201c [Greece], ZMHB (Berlin).-Synonymy in Tkalcu (1974a: 122).<\/p>\n

Distribution-Europe:<\/strong> BG, GR, GR(Aegean Islands), MK. <\/em>Northern Africa:<\/strong> ET<\/u>. <\/em>Southwestern Asia:<\/strong> IL, IRQ, JOR, RL, SYR, TR.<\/p>\n

Identification-Keys, Descriptions: <\/strong>Ducke (1900); Mavromoustakis (1948b: 216, 1959: 297-298); Tkalcu (1974a); M\u00fcller (2018a).<\/p>\n

Nesting biology-Nesting site: <\/strong>Preexisting cavities: empty snail shells of medium size (e.g. Theba<\/em>). Nesting material:<\/strong> Cell partitions and nest plug are made of chewed leaves. (M\u00fcller, 2018a and references therein)<\/p>\n

Flower preferences: <\/strong>Oligolectic on Asteraceae with a near exclusive preference for Carduoideae; species of the Cichorioideae and Asteroideae are very rarely exploited as pollen sources as well (M\u00fcller, 2018a).<\/p>\n

Osmia (Hoplosmia) spinulosa<\/em> (Kirby, 1802)<\/h3>\n

1802 Apis <\/em>spinulosa<\/em><\/strong> Kirby, Monographia Apum Angliae, 2: 261. Type material<\/u>: Lectotype m, by designation of Tkalcu (1974a: 119), [United Kingdom], BMNH (London); paralectotypes m, ff. Type species<\/u> of Hoplosmia<\/em> Thomson.-Combination Pseudosmia spinulosa<\/em> (Kirby) in Radoszkowski (1874c: 159). Combination Anthocopa (Hoplosmia) spinulosa<\/em> (Kirby) in Tkalcu (1974a: 119). Combination Hoplosmia (Hoplosmia) spinulosa<\/em> (Kirby) in Ungricht et al. (2008: 131).<\/p>\n

1882 Osmia<\/em> euchreiformis<\/em><\/strong> Radoszkowski, Proceedings of the Warsaw Society of Naturalists, Bulletin of Biology, 2: 77 [not seen]. Type material<\/u>: Holotype m, \u201eEchmiadzin\u201c [Armenia], ZMHB (Berlin).-Synonymy in Tkalcu (1974a: 119).<\/p>\n

Distribution-Europe: <\/strong>A, AND, ARM, B, BG, BIH, CH, CZ, D, DK, E, F, FL, GB<\/u>, GE, H, HR, I, I(Sa), I(Si), L, MD, MK, N, NL, PL, RO, RUS(CR,SR), S, SCG, SK, SLO, UA, UA(Crimea). Northern Asia:<\/strong> KS, KZ, RU(WS,ES), RUS(WS), TJ. Southwestern Asia:<\/strong> TR.<\/p>\n

Identification-Keys, Descriptions: <\/strong>Amiet et al. (2004); Banaszak and Romasenko (2001); Benoist (1931); Ducke (1900); Mavromoustakis (1959: 298); Medvedeva (1978); M\u00f3cz\u00e1r (1958); Radoszkowski (1874c: 159, 1887a: 282-283); Scheuchl (1996); Schmiedeknecht (1885-1886); Tkalcu (1974a, 1992: 220-221); M\u00fcller (2018a).<\/p>\n

Nesting biology-Nesting site: <\/strong>Preexisting cavities: empty snail shells of small to medium size (e.g. Cepaea, Cornuella<\/em>, Fruticicola, Helicella, Pomatias, Xerolenta, Zebrina, <\/em>small shells of Helix<\/em>) with 1-3, mostly 2 cells per shell. The nests are not sealed against their rear end with a basal wall and the outermost cell partition is distinctly more robust than all the other cell partitions and the nest plug, probably acting as a barrier against parasites and predators. The closed shells are not transported but turned so that the shell opening is directed towards the ground. Nesting material: <\/strong>Cell partitions and nest plug are made of chewed leaves (e.g. from Potentilla, Sanguisorba<\/em>). The shell surface is not plastered with patches of leaf pulp. (M\u00fcller, 2018a and references therein). Brood parasites:<\/strong> Stelis odontopyga<\/em> (Hymenoptera, Megachilidae), Chrysura cuprea<\/em> and C. trimaculata<\/em> (Hymenoptera, Chrysididae), Melittobia acasta<\/em> (Hymenoptera, Eulophidae), Pteromalus apum<\/em> and P. venustus<\/em> (Hymenoptera, Pteromalidae) and Anthrax aethiops<\/em> (Diptera, Bombyliidae) (Bl\u00fcthgen, 1926; Noskiewicz, 1926; Kunz, 1994; M\u00fcller, 1994; BWARS, 2013).<\/p>\n

Flower preferences: <\/strong>Oligolectic on Asteraceae; pollen host plants include members of the Carduoideae, the Cichorioideae and the Asteroideae (M\u00fcller, 2018a).<\/p>\n

Osmia (Hoplosmia) tyrneri<\/em> (Tkalcu, 1992)<\/h3>\n

1992 Hoplosmia (Hoplosmia) <\/em>tyrneri<\/em><\/strong> Tkalcu, Acta Societatis Zoologicae Bohemoslovacae (Praha), 56: 220. Type material<\/u>: Holotype m, \u201eTashkent, Galvasai\u201c [Uzbekistan], Tyrner Collection (Litvinov); paratypes mm.<\/p>\n

Distribution-Northern Asia:<\/strong> KZ, UZ<\/u>.<\/em><\/p>\n

Identification-Keys, Descriptions: <\/strong>M\u00fcller (2018a).<\/p>\n

Nesting biology: <\/strong>Oligolectic on Asteraceae; pollen host plants include members of the Cichorioideae and more rarely of the Carduoideae and the Asteroideae (A. M\u00fcller, unpublished, based on 15 pollen loads from 2 different localities).<\/p>\n

Flower preferences: <\/strong>Unknown.<\/p>\n

bidentata<\/u><\/em> species group<\/u><\/h3>\n

Osmia (Hoplosmia) anceyi<\/em> P\u00e9rez, 1879<\/h3>\n

1879 Osmia <\/em>Anceyi<\/em><\/strong> P\u00e9rez, Actes de la Soci\u00e9t\u00e9 Linn\u00e9enne de Bordeaux, 33: 187. Type material<\/u>: Lectotype f, by designation of Tkalcu (1974a: 128), \u201eEnvirons de Marseille\u201c [France], MNHN (Paris).- Combination Anthocopa (Odontanthocopa) anceyi<\/em> (P\u00e9rez) in Tkalcu (1974a: 128). Combination Hoplosmia (Odontanthocopa) anceyi<\/em> (P\u00e9rez) in Ungricht et al. (2008: 132).<\/p>\n

Distribution-Europe:<\/strong> CH, E, F<\/u>, F(Co), P. Northern Africa:<\/strong> DZ, MA, TN.<\/em><\/p>\n

Identification-Keys, Descriptions: <\/strong>Amiet et al. (2004); Benoist (1931); Schmiedeknecht (1885-1886); Tkalcu (1974a); M\u00fcller (2018a).<\/p>\n

Nesting biology-Nesting site: <\/strong>Preexisting cavities: insect burrows in dead wood, dead hollow plant stems, burrows in the pith of dead stems (e.g. Asphodelus<\/em>) excavated by other aculeates (e.g. Ceratina<\/em>) or abandoned nests of aculeate hymenopterans in clay walls. The nests contain up to nine linearly arranged brood cells. Nesting material: <\/strong>Cell partitions and nest plug are made of mud, sometimes mixed with particles of pith. (M\u00fcller, 2018a and references therein)<\/p>\n

Flower preferences: <\/strong>Oligolectic on Asteraceae with a probably exclusive preference for Carduoideae (M\u00fcller, 2018a).<\/p>\n

– <\/em>subspecies anceyi<\/em><\/strong> P\u00e9rez, 1879<\/h3>\n

1879 Automatically established nominotypical subspecific taxon (original description and type specimen are identical with those of the species rank taxon of the same name above).<\/p>\n

Distribution-Europe:<\/strong> CH, E, F<\/u>, P.<\/p>\n

Identification-Keys, Descriptions: <\/strong>M\u00fcller (2018a).<\/p>\n

– <\/em>subspecies biarmica<\/em><\/strong> (Zanden, 1994)<\/h3>\n

1994 Hoplosmia (Odontanthocopa) anceyi <\/em>biarmica<\/em><\/strong> Zanden, Linzer Biologische Beitr\u00e4ge, 26: 1114. Type material<\/u>: Holotype f, \u201eTizi-n-Tichka S., 2000 m\u201c [Morocco], Schwarz Collection (Ansfelden); paratypes mm, ff.<\/p>\n

Distribution-Northern Africa:<\/strong> DZ, MA<\/u>, TN.<\/em><\/p>\n

Identification-Keys, Descriptions: <\/strong>M\u00fcller (2018a).<\/p>\n

Osmia (Hoplosmia) bidentata<\/em> Morawitz, 1875<\/h3>\n

1875 Osmia <\/em>bidentata<\/em><\/strong> Morawitz, Horae Societatis Entomologicae Rossicae (St. Petersburg), 12: 38. Type material<\/u>: Syntypes m(m), f(f), \u201eEtschmiadzin\u201c [Armenia]. Type species<\/u> of Odontanthocopa<\/em> Tkalcu.-Combination Anthocopa (Odontanthocopa) bidentata<\/em> (Morawitz) in Tkalcu (1974a: 127). Combination Hoplosmia (Odontanthocopa) bidentata<\/em> (Morawitz) in Ungricht et al. (2008: 133).<\/p>\n

1877 Osmia<\/em> affinis<\/em><\/strong> Frivaldsky, Mathematikai \u00e9s Term\u00e9szettudom\u00e1nyi K\u00f6zlem\u00e9nyek, 13: 360. Type material<\/u>: Syntypes m(m), f(f), \u201eIn Comitatu Crassoviensi ad Oraviczam, in Comitatu Pestiensi vero ad Budapestinum et in Praedio-Pesz\u00e9r\u201c [Hungary].-Synonymy in Tkalcu (1974a: 127).<\/p>\n

Distribution-Europe:<\/strong> A, AL, ARM<\/u>, AZ, BG, CZ, F, GR, GR(Aegean Islands, Cr), H, HR, I, MK, PL, RO, RUS(CR,SR), SCG, SK, SLO, UA, UA(Crimea). <\/em>Northern Africa:<\/strong> ET. <\/em>Southwestern Asia:<\/strong> IL, IR, IRQ, JOR, RL, SYR, TR.<\/em><\/p>\n

Identification-Keys, Descriptions: <\/strong>Alfken (1914: 197); Banaszak and Romasenko (2001); Benoist (1931); Ducke (1900); Medvedeva (1978); M\u00f3cz\u00e1r (1958); Romankova (1995); Scheuchl (1996); Schmiedeknecht (1885-1886); Tkalcu (1974a, 1992: 225); M\u00fcller (2018a).<\/p>\n

Nesting biology-Nesting site: <\/strong>Preexisting cavities: insect burrows in dead wood, hollow stems (e.g. Phragmites<\/em>) or cavities in the soil, possibly also abandoned nests of aculeates in loess scarps. The nests contain 1-11 linearly arranged brood cells. Nesting material: <\/strong>Cell partitions and nest plug are made of mud. (M\u00fcller, 2018a and references therein; Ivanov et al., 2019)<\/p>\n

Flower preferences: <\/strong>Oligolectic on Asteraceae with a probably exclusive preference for Carduoideae (A. M\u00fcller, 2018a). In contrast, G\u00fcler & Sorkun (2007) assume O. bidentata<\/em> to be polylectic, harvesting pollen from eleven plant families.<\/p>\n

– <\/em>subspecies bidentata<\/em><\/strong> Morawitz, 1875<\/h3>\n

1875 Automatically established nominotypical subspecific taxon (original description and type specimen are identical with those of the species rank taxon of the same name above).<\/p>\n

Distribution-Europe:<\/strong> A, AL, ARM<\/u>, BG, CZ, GR, GR(Cr), H, HR, I, MK, PL, RO, RUS(CR,SR), SCG, SK, SLO, UA, UA(Crimea). <\/em><\/p>\n

Identification-Keys, Descriptions: <\/strong>Tkalcu (1979: 320); M\u00fcller (2018a).<\/p>\n

– <\/em>subspecies pallens<\/em><\/strong> (Tkalcu, 1979)<\/h3>\n

1979 Anthocopa (Odontanthocopa) bidentata <\/em>pallens<\/em><\/strong> Tkalcu, Acta Entomologica Bohemoslovaca (Praha), 76: 320. Type material<\/u>: Holotype f, \u201eErzurum\u201c [Turkey], EMET (Erzurum); paratypes mm, ff.-Combination Hoplosmia (Odontanthocopa) bidentata pallens<\/em> (Tkalcu) in Ungricht et al. (2008: 133).<\/p>\n

Distribution-Europe: <\/strong>AZ, GR(Lesbos). Northern Africa:<\/strong> ET. <\/em>Southwestern Asia:<\/strong> IL, IR, IRQ, JOR, SYR, TR<\/u>.<\/em><\/p>\n

Identification-Keys, Descriptions: <\/strong>M\u00fcller (2018a).<\/p>\n

ligurica<\/u><\/em> species group<\/u><\/h3>\n

Osmia (Hoplosmia) dido <\/em>Gribodo, 1894<\/h3>\n

1894 Osmia <\/em>dido<\/em><\/strong> Gribodo, Bullettino della Societ\u00e0 Entomologica Italiana, 26: 289. Type material<\/u>: Holotype f, \u201eAlgeria\u201c [Algeria], MSNG (Genova).-Combination Anthocopa dido <\/em>(Gribodo in Zanden (1990: 54). Combination Hoplosmia (Odontanthocopa) dido<\/em> (Gribodo) in Ungricht et al. (2008: 133).<\/p>\n

1895 Osmia<\/em> abbreviata<\/em><\/strong> P\u00e9rez, Esp\u00e8ces nouvelles de mellif\u00e8res de Barbarie, p. 13. NOMEN PRAEOCCUPATUM [not Osmia abbreviata <\/em>Morawitz, 1875]. Type material<\/u>: Lectotype f, by designation of Tkalcu (1974a: 128), \u201eAlg\u00e9ria\u201c [Algeria], MNHN (Paris).-Synonymy in Zanden (1990: 54).<\/p>\n

1896 Osmia <\/em>compacta<\/em><\/strong> P\u00e9rez, Correctures: Esp\u00e8ces nouvelles de mellif\u00e8res de Barbarie, not paginated single page appendix to P\u00e9rez (1895). NOMEN NOVUM with same type specimen for preoccupied Osmia abbreviata<\/em> P\u00e9rez, 1895 [not Osmia abbreviata<\/em> Morawitz, 1875].-Combination Anthocopa (Odontanthocopa) compacta<\/em> (P\u00e9rez) in Tkalcu (1974a: 128).<\/p>\n

Distribution-Northern Africa:<\/strong> DZ<\/u>, MA, TN. <\/em><\/p>\n

Identification-Keys, Descriptions: <\/strong>Alfken (1914: 197); Tkalcu (1974a, 1979: 320-321); M\u00fcller (2018a).<\/p>\n

Nesting biology: <\/strong>Unknown.<\/p>\n

Flower preferences: <\/strong>Probably oligolectic on Asteraceae with a strong or even exclusive preference for Carduoideae (M\u00fcller, 2018a).<\/p>\n

Osmia (Hoplosmia) distinguenda<\/em> (Tkalcu, 1974)<\/h3>\n

1974 Anthocopa (Odontanthocopa) <\/em>distinguenda<\/em><\/strong> Tkalcu, Acta Entomologica Bohemoslovaca (Praha), 71: 129. Type material<\/u>: Holotype m, \u201eJerusalem\u201c [Israel], BMNH (London).-Combination Hoplosmia (Odontanthocopa) distinguenda<\/em> (Tkalcu) in Ungricht et al. (2008: 134).<\/p>\n

Distribution-Europe:<\/strong> ARM, AZ, GR. <\/em>Southwestern Asia:<\/strong> IL<\/u>, IRQ, JOR, RL, SYR, TR.<\/em><\/p>\n

Identification-Keys, Descriptions: <\/strong>Tkalcu (1979: 320-321, 1992: 223-224).<\/p>\n

Nesting biology-Nesting site: Preexisting cavities: small cavities in rocks and stones. Nesting material: Brood cells entirely or partially built from chewed leaves, nest plug also made of leaf pulp. (M\u00fcller, 2018a and references therein)<\/p>\n

Flower preferences: <\/strong>Oligolectic on Asteraceae with a strong or even exclusive preference for Carduoideae (A. M\u00fcller, 2018a).<\/p>\n

Osmia (Hoplosmia) hermonensis<\/em> (Tkalcu, 1992)<\/h3>\n

1992 Hoplosmia (Odontanthocopa) <\/em>hermonensis<\/em><\/strong> Tkalcu, Acta Societatis Zoologicae Bohemoslovacae (Praha), 56: 223. Type material<\/u>: Holotype m, \u201eMt. Hermon, 1700 m\u201c [Israel], RMNH (Leiden); paratypes mm.<\/p>\n

Distribution-Southwestern Asia:<\/strong> IL<\/u>.<\/em><\/p>\n

Identification-Keys, Descriptions: <\/strong>M\u00fcller (2018a). Note:<\/strong> Female unknown.<\/p>\n

Nesting biology: <\/strong>Unknown.<\/p>\n

Flower preferences: <\/strong>Unknown.<\/p>\n

Osmia (Hoplosmia) larochei<\/em> Tkalcu, 1993<\/h3>\n

1993 Osmia (Odontanthocopa) <\/em>larochei<\/em><\/strong> Tkalcu, Ver\u00f6ffentlichungen des \u00dcbersee-Museum Bremen (Naturwissenschaften), 12: 810. Type material<\/u>: Holotype f, \u201eGran Canaria Cueva Grande Las Palmas\u201c [Spain: Canary Islands], La Roche Collection; paratype f.-Combination Hoplosmia (Odontanthocopa) larochei<\/em> (Tkalcu) in Ungricht et al. (2008: 134).<\/p>\n

Distribution-Northern Africa:<\/strong> E(Ca)<\/u>.<\/em><\/p>\n

Identification-Keys, Descriptions: <\/strong>Tkalcu (2000a: 310); M\u00fcller (2018a).<\/p>\n

Nesting biology: <\/strong>Unknown.<\/p>\n

Flower preferences: <\/strong>Unknown. Flower record: Andryala pinnatifolia<\/em> (Asteraceae) (Hohmann et al., 1993).<\/p>\n

Osmia (Hoplosmia) ligurica<\/em> Morawitz, 1868<\/h3>\n

1868 Osmia <\/em>ligurica<\/em><\/strong> Morawitz, Horae Societatis Entomologicae Rossicae (St. Petersburg), 5: 150. Type material<\/u>: Syntypes m(m), f(f), \u201eBei Nizza im Thale des Paglione\u201c [France].-Combination Anthocopa (Odontanthocopa) ligurica<\/em> (Morawitz) in Tkalcu (1974a: 130). Combination Hoplosmia (Odontanthocopa) ligurica<\/em> (Morawitz) in Ungricht et al. (2008: 135).<\/p>\n

1879 Osmia<\/em> detrita<\/em><\/strong> P\u00e9rez, Actes de la Soci\u00e9t\u00e9 Linn\u00e9enne de Bordeaux, 33: 188. Type material<\/u>: Lectotype f, by designation of Tkalcu (1974a: 130), \u201eBordeaux, Marseille\u201c [France], MNHN (Paris).-Synonymy in Schmiedeknecht (1886: 199 [1065]).<\/p>\n

Distribution-Europe:<\/strong> A, AL, ARM, AZ, BG, CH, CZ, E, E(Ba), F<\/u>, F(Co), GE, GR, GR(Aegean Islands, Cr), H, HR, I, I(Sa), I(Si), P, MK, RO, RUS(SR), SCG, SK, SLO. <\/em>Northern Africa: <\/strong>MA. Northern Asia:<\/strong> TM. Southwestern Asia:<\/strong> CY, IL, IR, IRQ, JOR, SYR, TR.<\/em><\/p>\n

Identification-Keys, Descriptions: <\/strong>Amiet et al. (2004); Banaszak and Romasenko (2001); Benoist (1931); Ducke (1900); Medvedeva (1978); M\u00f3cz\u00e1r (1958); Schmiedeknecht (1885-1886); Tkalcu (1974a, 1992: 223); M\u00fcller (2018a).<\/p>\n

Nesting biology-Nesting site: <\/strong>Preexisting cavities: dead hollow stems or burrows in pithy stems (e.g. Artemisia, Foeniculum, Rubus, Scolymus<\/em>) excavated by other aculeates (e.g. Ceratina<\/em>). The nests contain up to 8 linearly arranged brood cells. Nesting material: <\/strong>Cell partitions and nest plug are made of chewed leaves. (M\u00fcller, 2018a and references therein)<\/p>\n

Flower preferences: <\/strong>Oligolectic on Asteraceae with a preference for Asteroideae; species of the Cichorioideae are also exploited (M\u00fcller, 2018a).<\/p>\n

Osmia (Hoplosmia) olgae<\/em> (Tkalcu, 1978)<\/h3>\n

1978 Anthocopa (Odontanthocopa) <\/em>olgae<\/em><\/strong> Tkalcu, Casopis Slezskeho Muzea Opava (A), 27: 159. Type material<\/u>: Holotype m, \u201eBulgaria sept.-or. Albena\u201c [Bulgaria], Tkalcu Collection (Praha); paratypes mm, ff.-Combination Hoplosmia (Odontanthocopa) olgae<\/em> (Tkalcu) in Ungricht et al. (2008: 135).<\/p>\n

Distribution-Europe: <\/strong>ARM, BG<\/u>. <\/em>Southwestern Asia:<\/strong> TR.<\/em><\/p>\n

Identification-Keys, Descriptions: <\/strong>M\u00fcller (2018a).<\/p>\n

Nesting biology: <\/strong>Unknown.<\/p>\n

Flower preferences: <\/strong>Unknown. The only two pollen loads analyzed so far consisted of pollen of Asteroideae (Asteraceae) (M\u00fcller, 2018a).<\/p>\n

Osmia (Hoplosmia) padri<\/em> (Tkalcu, 1974)<\/h3>\n

1974 Anthocopa (Odontanthocopa) <\/em>padri<\/em><\/strong> Tkalcu, Acta Entomologica Bohemoslovaca (Praha), 71: 129. Type material<\/u>: Holotype f, \u201eSlancev Brjag\u201c [Bulgaria], P\u00e1dr Collection (Praha); paratypes mm, ff.-Combination Hoplosmia (Odontanthocopa) padri<\/em> (Tkalcu) in Ungricht et al. (2008: 135).<\/p>\n

Distribution-Europe:<\/strong> BG<\/u>, GR, GR(Aegean Islands), HR, MK, SCG.<\/em> Southwestern Asia:<\/strong> IL, IRQ, SYR, TR.<\/em><\/p>\n

Identification-Keys, Descriptions: <\/strong>Tkalcu (1979: 321); M\u00fcller (2018a).<\/p>\n

Nesting biology-Nesting site: <\/strong>Preexisting cavities: insect burrows in dead wood. Nesting material: <\/strong>Unknown. (M\u00fcller, 2018a and references therein)<\/p>\n

Flower preferences: <\/strong>Oligolectic on Asteraceae with a probably exclusive preference for Carduoideae (M\u00fcller, 2018a).<\/p>\n

Osmia (Hoplosmia) scutellaris<\/em> Morawitz, 1868<\/h3>\n

1868 Osmia <\/em>scutellaris<\/em><\/strong> Morawitz, Horae Societatis Entomologicae Rossicae (St. Petersburg), 5: 151. Type material<\/u>: Syntypes m(m), f(f), \u201eim Magnan-Thale\u201c [France], \u201eauch n\u00f6rdlicher bei Pallanza\u201c [Italy], \u201eund Lugano\u201c [Switzerland].-Combination Anthocopa (Odontanthocopa) scutellaris<\/em> (Morawitz in Tkalcu (1974a: 130). Combination Hoplosmia (Odontanthocopa) scutellaris <\/em>(Morawitz) in Ungricht et al. (2008: 135).<\/p>\n

1934 Heriades <\/em>integra<\/em><\/strong> Benoist, Bulletin de la Soci\u00e9t\u00e9 Entomologique de France, 39: 159. Type material<\/u>: Holotype m, \u201eTanger\u201c [Morocco], MNHN (Paris).-Combination Stenoheriades integer<\/em> (Benoist) in Griswold (1994) and Ungricht et al. (2008). Synonymy in M\u00fcller & Trunz (2014).<\/p>\n

Distribution-Europe:<\/strong> AL, AND, ARM, AZ, BG, CH<\/u>, E, F<\/u>, F(Co), GE, GR, GR(Aegean Islands, Cr), H, HR, I<\/u>, I(Sa), I(Si), MK, P, RO, RUS(SR), SCG, SK, SLO, UA, UA(Crimea). <\/em>Northern Africa: <\/strong>DZ, MA, TN. Northern Asia: <\/strong>TM. Southwestern Asia:<\/strong> CY, IL, IR, IRQ, JOR, RL, SYR, TR.<\/em><\/p>\n

Identification-Keys, Descriptions: <\/strong>Amiet et al. (2004); Banaszak and Romasenko (2001); Benoist (1931); Ducke (1900); Medvedeva (1978); Schmiedeknecht (1885-1886); Tkalcu (1974b, 1978: 159-162, 1993b: 810-811, 2000a: 310); M\u00fcller (2018a).<\/p>\n

Nesting biology-Nesting site: <\/strong>Preexisting cavities: dead hollow stems or burrows in pithy stems (e.g. Rubus<\/em>) excavated by other aculeates (e.g. Ceratina<\/em>). Nesting material: <\/strong>Unknown. (M\u00fcller, 2018a and references therein)<\/p>\n

Flower preferences: <\/strong>Oligolectic on Asteraceae with a preference for Cichorioideae; species of the Asteroideae are also exploited (M\u00fcller, 2018a).<\/p>\n

Osmia (Hoplosmia) warnckei<\/em> (Tkalcu, 1992)<\/h3>\n

1992 Hoplosmia (Odontanthocopa) <\/em>warnckei<\/em><\/strong> Tkalcu, Acta Societatis Zoologicae Bohemoslovacae (Praha), 56: 223. Type material<\/u>: Holotype m, \u201e15 km SE Sarvestan\/Fars, 1800 m\u201c [Iran], OLML (Linz); paratypes mm.<\/p>\n

Distribution-Southwestern Asia:<\/strong> IR<\/u>, IRQ, TR.<\/em><\/p>\n

Identification-Keys, Descriptions: <\/strong>M\u00fcller (2018a).<\/p>\n

Nesting biology: <\/strong>Unknown.<\/p>\n

Flower preferences: Unknown. <\/strong>The only pollen load analyzed so far consisted of pollen of Carduoideae (Asteraceae) (M\u00fcller, 2018a).<\/p>\n","protected":false},"excerpt":{"rendered":"

The subgenus Hoplosmia Thomson is confined to the Palaearctic region. It contains 21 species (M\u00fcller, 2018a). Female of Osmia (Hoplosmia) spinulosa (Kirby, 1802) collecting pollen on a flower head of Buphthalmum salicifolium (Asteraceae). Foto A. M\u00fcller. Species accounts spinulosa species group Osmia (Hoplosmia) carbo (Zanden, 1994) 1994 Hoplosmia (Paranthocopa) pinguis carbo Zanden, Linzer Biologische Beitr\u00e4ge, […]<\/p>\n","protected":false},"author":1092,"featured_media":0,"parent":46,"menu_order":0,"comment_status":"open","ping_status":"closed","template":"","meta":{"footnotes":""},"class_list":["post-144","page","type-page","status-publish","hentry"],"_links":{"self":[{"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/pages\/144","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/pages"}],"about":[{"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/types\/page"}],"author":[{"embeddable":true,"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/users\/1092"}],"replies":[{"embeddable":true,"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/comments?post=144"}],"version-history":[{"count":5,"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/pages\/144\/revisions"}],"predecessor-version":[{"id":7433,"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/pages\/144\/revisions\/7433"}],"up":[{"embeddable":true,"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/pages\/46"}],"wp:attachment":[{"href":"https:\/\/blogs.ethz.ch\/osmiini\/wp-json\/wp\/v2\/media?parent=144"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}